Publications (2008 - present)

Preprints available online

Metz, J., Lampei, C., Bäumler, L., Bocherens, H., Dittberner, H., Hennerberg, L., de Meaux, J., Tielbörger, K., preprint. Rapid adaptive evolution to drought in a subset of traits in a multi-site climate change experiment. Authorea doi:

In Press

Bocherens, H., Drucker, D.G., in press. Isotopic insights on the ecological interactions between humans and woolly mammoths during the Middle and Upper Palaeolithic in Europe. Proceedings of the Human-Elephants congress, University of Tübingen Press.

Díaz-Zorita Bonilla, M., Waterman, A.J., Peate, D. in press. La movilidad de las poblaciones de la Edad del Cobre en el Suroeste de España a través de los análisis de isótopos estables de 87Sr/86Sr (Human mobility during the Copper and Bronze Age through isotopical analysis of 87Sr/86Sr). Proceedings of the 2nd Conference of Prehistory of Andalusia. February 15th-17th, 2012, Antequera, Málaga, Spain.

Germonpré, M., Bocherens, H., Bessudnov, A., Lázničková-Galetová, M., Reynolds, N., Sablin, M., Wißing, C., in press. Seasonality at Middle and Upper Palaeolithic sites based on the presence and wear of molars from nursing mammoth calves. Proceedings of the Human-Elephants congress, University of Tübingen Press.

Germonpré, M., Láznickova-Galetova, M., Sablin, M., Bocherens, H., in press. 7. Could incipient dogs have enhanced diferential access to resources among Upper Palaeolithic hunter-gatherers in Europe? In (L. Moreau Ed.): Social inequality before farming? Muldisciplinary approaches to the investigation of egalitarian and non-egalitarian social relationships in prehistoric and extant hunter-gatherer societies. McDonald Institute of Achaeology, Cambridge.

Germonpré, M., Láznickova-Galetova, M., Sablin, M., Bocherens, H., in press. The advantages of owning a Palaeolithic dog. In: Dogs Past and Present Archaeopress.

* Habinger, S., De Cupere, B., Dövener, F., Pucher, E., Bocherens, H. in press. Mobility and origin of camels in the Roman Empire through serial stable carbon and oxygen isotopes variations in tooth enamel. Quaternary International

* Heidgen, S., Marinova, E., Krauß, R., Nelle, O., Ebner, M., Märkle, T., Miranda, T., Bofinger, J., Klingler, S., Junginger, A., in press. Palaeoenvironment and potential resources for early holocene subsistence in the Ammer River Valley (Germany) based on palaeoecological and bioarchaeological evidence. Quaternary International

Lord, E., Dussex, N., Kierczak, M., Díez-del-Molino, D., Ryder, O.A., Stanton, D.W.G., Gilbert, M.T.P., Sánchez-Barreiro F., Zhang G., Sinding, M.H.S., Lorenzen E.D., Fedorov S., Bocherens, H., Nathan, S., Goosens, B., van der Plicht, J., Chan, Y.L., Prost, S., Potapova O., Kirillova, I., Lister, A.M., Heintzman, P.D., Kapp, J., Shapiro, B., Vartanyan, S., Götherström, A., Dalén, L., 2020. Pre-extinction demographic stability and genomic signatures of adaptation in the woolly rhinoceros. Current Biology 30: 1-9. media report 1 media report 2 media report 3

* Metz, J., Lampei, C., Bäumler, L., Bocherens, H., Dittberner, H., Henneberg, L., de Meaux, J., Tielbörger, K., in press. Rapid adaptive evolution to drought in a subset of traits in a multi-site climate change experiment. Ecology Letters

* Miranda, T., Roth-Nebelsick, A., Junginger, A., Ebner, M., in press. Habitat conditions, spatial districution and trichome morphology of different species of Tillandsia growing on trees on the Ilha Grande Island, Brazil. Flora 

* Rufí, I., Drucker, D.G., Bocherens. H., Lloveras, L., Soler, J., Soler, N., in press. Revision on the occurrence of musk ox (Ovibos moschatus Zimmermann 1780) from the Gravettian of Arbreda Cave (Serinyà, Northeast of Iberian Peninsula): New insights for the study of Iberian cold-adapted faunas. Boreas

Suraprasit, K., Jaeger, J.-J., Chaimanee, Y., and Sucharit, C. in press. Taxonomic reassessment of large mammals from the Pleistocene Homo-bearing site of Tham Wiman Nakin (Northeast Thailand): relevance for faunal patterns in mainland Southeast Asia. Quaternary International 


* Allabar, A., Dobson, K.J., Bauer, C.C., Nowak, M., 2020. Vesicle shrinkage in hydrous phonolitic melt during cooling. Contributions to Mineralogy and Petrology 175: 21. 

* Baumann, C., Bocherens, H., Drucker, D.G., Conard, N.J., 2020. Fox dietary ecology as a tracer of human impact on Pleistocene ecosystems. PLoS ONE 15(7): e0235692.    

* Baumann, C., Starkovich, B.M., Drucker, D.G., Münzel, S.C., Conard, N.C., Bocherens, H. 2020. Dietary niche partitioning among Magdalenian canids in southwestern Germany and Switzerland. Quaternary Science Reviews 227: 106032. 

* Dantas, M.A.T., Cherkinsky, A., Lessa, C.M.F., Santos, L.V., Cozzuol, M.A., Omena, É.C., Lopes da Silva, J.L., Sial, A.N., Bocherens, H., 2020. Isotopic paleoecology (δ13C, δ18O), of a Late Pleistocene vertebrate community from Sergipe, NE Brazil. Revista Brasileira de Paleontologia 23(2): 138-152. https://doi.10.4072/rbp.2020.2.05 

Drucker, D.G., Bridault, A., Ducrocq, T., Baumann, C., Valentin, F., 2020. Environment and human subsistence in Northern France at the Late Glacial to early Holocene transition. Archaeological and Anthropological Sciences, 12(8): 1-16. 

Ebner, M., Poli, F., Keil, R., Radtke, R., Miranda, T., 2020. In: (Werneburg, I., Betz, O., Eds) Phylogenie, Funktionsmorphologie und Bionik. Schriften zum 60. Phylogenetischen Symposium in Tübingen. Scidinge Hall Verlag Tübingen, 332 p.

Elalouf, J.-M., Bocherens, H., Bon, C., Fosse, Ph., Palacio, P., Philippe, M., 2020. 6.4. Restes osseux et analyse ADN: apports paléoenvironnementaux. In: Atlas de la grotte Chauvet-Pont d’Arc, Delannoy J.J. & Geneste J.M. (Eds), Collection “Documents d’archéologie française”, Editions Fondation Maison des Sciences de l’Homme, pp. 150-152.

* Fuentes, R., Ono, R., Carlos, J., Kerfant, C., Sriwigati, Miranda, T., Aziz, N., Sofian, H.O., Pawlik, A., 2020. Stuck with notches: Direct evidence of plant processing during the last glacial maximum to Holocene in North Sulawesi. Journal of Archaeological Science: Reports 30: 102207. 

* Grohé, C., de Bonis, L., Chaimanee, Y., Chavasseau, O., Rugbumrung, M., Yamee, C., Suraprasit, K., Gibert, C., Surault, J., Blondel, C., Jaeger, J.-J., 2020. The late middle Miocene Mae Moh Basin of northern Thailand: the richest Neogene assemblage of Carnivora from Southeast Asia and a paleobiogeographic analysis of Miocene Asian carnivorans. American Museum Novitates 3957: 1-57. pdf

* Krajcarz, M., Krajcarz, M.T., Baca, M., Baumann. C., van Neer, W., Popović, D., Sudoł-Procyk, M., Wach, B., Wilczyński, J., Wojenka, M., Bocherens, H., 2020. Ancestors of domestic cats in Neolithic Central Europe – isotopic evidence of a synanthropic diet. PNAS 117(30): 17710-17719.   media report 1 media report 2

* Lipson, M., Ribot, I., Mallick, S., Rohland, N., Olalde, I., Adamski, N., Broomandkhoshbacht, N., Lawson, A.M., Lopez, S., Oppenheimer, J., Stewardson, K., Asombang, R.. Bocherens, H., Bradman, N., Cornelissen, E., Crevecoeur, I.. Lavachery, P.. de Maret, P., Mindzie, C.M., Orban, R., Swachuk, E., Semal, P., Patterson, N., Hellenthal, G., Lalueza-Fox, C., MacEachern, S., Prendergast, M., Reich, D., 2020. Ancient West African foragers in the context of deep human genetic history. Nature 577: 665-670. media report 1 media report 2 media report 3

* Lister, A., Rivals, F., Barkai, R., Bocherens, H., 2020. Current research on mammoths and their relatives. Quaternary Science Reviews 231: 106174. 

* Loog, L., Thalmann, O., Sinding, M.-H.S., Schuenemann, V., Perri, A., Germonpré, M., Bocherens, H., Witt, K.E., Samaniego Castruita, J.A., Velasco, M.S., Lundstrøm, I.K.C., Wales, N., Sonet, G.E., Frantz, L., Schroeder, H., Budd, J., Elodie-Laure, J., Fedorov, S., Gasparyan, B., Kandel, A.W., Lázničková-Galetová, M., Napierala, H., Uerpmann, H.-P., Nikolskiy, P.A., Pavlova, E.Y., Pitulko, V.V., Herzog, K.-H., Malhi, R.S., Willerslev, E., Hansen, A.J., Dobney, K., Gilbert, T.P., Krause, J., Larsson G., Eriksson, A., Manica, A., 2020. Modern wolves trace their ancestry to a late Pleistocene expansion from Beringia. Molecular Ecology 162: 2015-2025. 

* Meadows, J., Lozovskaya, O., Bondetti, M., Drucker, D.G., Moiseyev, V., 2020. Human palaeodiet at Zamostje 2, central Russia: Results of radiocarbon and stable isotope analysis. Quaternary International 541: 89-103. 

Meleg, I.N., Naito, Y.I., Bocherens, H., 2020. Herbivory in Carnivora: the case of emblematic cave bears. Behind the article (Nature Research).

* Naito, Y., Meleg. I., Robu, M., Drucker, D.G., Wißing, C., Hofreiter, M., Barlow, A., Bocherens, H., 2020. Heavy reliance on plants for Romanian cave bears evidenced by amino acid nitrogen isotope analysis. Scientific Reports 10: 6612. media report 1, media report 2 TV interview

* Oinonen, M., Alenius, T., Arppe, L., Bocherens, H., Etu-Sihvola, H., Helama, S., Huhtamaa, H., Lahtinen, M., Mannermaa, K., Onkamo, P., Vanhanen, S., Wessman, A. in press. Burdened by nature, buried in water - Resilience carried the Iron Age people through Fimbulwinter. PLoS ONE 15(4): e0231787.   

* Owocki, K., Kremer, B., Cotte, M., Bocherens, H., 2020. Diet preferences and climate inferred from Oxygen and Carbon isotopes of tooth enamel of Tarbosaurus bataar (Nemegt Formation, Mongolia). Palaeogeography, Palaeoclimatology, Palaeoecology 537: 109190. media report

* Pushkina, D., Saarinen, J., Ziegler, R., Bocherens, H., 2020. Comparisons in stable isotopic and mesowear paleoreconstructions of diet and habitat of the Middle and Late Pleistocene mammals in South-Western Germany. Quaternary Science Reviews 227: 106026.

* Ring, S.J., Bocherens, H., Wings, O., Rabi, M., 2020. Divergent mammalian body size in a stable Eocene greenhouse climate. Scientific Reports media report 1 media report 2

* Rivals, F., Drucker, D.G., Weber, M.-J., Audouze, F., Enloe, J.G., 2020. Dietary traits and habitats of the reindeer (Rangifer tarandus) during the Late Glacial of Northern Europe. Archaeological and Anthropological Sciences 12:98.

* Suraprasit, K., Jaeger, J.-J., Shoocongdej, R., Chaimanee, Y., Wattanapituksakul, A., Bocherens, H., 2020. Long-term isotope evidence on the diet and habitat breadth of Pleistocene to Holocene caprines in Thailand: implications for the extirpation and conservation of Himalayan gorals. Frontiers in Ecology and Evolution 8: article 67.

*Tschopp, E., Araújo, R., Brusatte, S.L., Hendrickx, C., Macaluso, L., Maidment, S.C.R., Rabi, M., Rashid, D., Romano, C., Williamson, T., 2020. Dinosaurs, but not only: Vertebrate evolution in the Mesozoic. In: E. Martinetto et al. (eds), Nature through time. Springer textbooks in Earth Sciences, Geography and Environment. 

Uhl, D., Bocherens, H., El Atfy, H., Havlik, P., Methner, K., Mulch, A., Solórzano Kraemer, M.M., 2020. Die Umwelt zu Edmonds Lebzeiten. Senckenberg / Natur-Forschung-Museum 150: 56-65. 

Valladas, H., Bocherens, H., Bon, C., Fosse, Ph., Philippe, M., Quiles, A., Geneste, J.-M., Monney, J., 2020. 5.4. Datation 14C d’ossements. In: Atlas de la grotte Chauvet-Pont d’Arc, Delannoy J.J. & Geneste J.M. (Eds), Collection “Documents d’archéologie française”, Editions Fondation Maison des Sciences de l’Homme, p. 125.

* Wong, G.L., Drucker, D.G., Starkovich, B.M., Conard, N.J., 2020.  Latest Pleistocene paleoenvironmental reconstructions from the Swabian Jura, southwestern Germany: Evidence from stable isotope analysis and micrommal remains. Palaeogeography, Palaeoclimatology, Palaeoecology 540: 109527.  

* Wong, G.L., Starkovich, B.M., Drucker, D.G., Conard, N.J., 2020. New perspectives on human subsistence during the Magdalenian in the Swabian Jura, Germany. Archaeological and Anthropological Sciences, 12(9): 1-31.


* Arppe, L., Karhu, J.A., Vartanyan, S., Drucker, D.G., Etu-Sihvola, H., Bocherens, H., 2019. Thriving or surviving? The isotopic record of the Wrangel Island woolly mammoth population. Quaternary Science Reviews 222: 105889.  media report 1 media report 2 media report 3 media report 4

* Bocherens, H. 2019. Isotopic insights on cave bear palaeodiet. Historical Biology 31: 410-421. doi: 10.1080/08912963.2018.14465419 media report

Bocherens, H., 2019.  Laudation: Dr. Frido Welker, twentieth recipient of the Tübingen Prize for Early Prehistory and Quaternary Ecology. Mitteilungen der Gesellschaft für Urgeschichte 27: 219-222.

Bocherens, H., 2019. Der Beginn der Anthroposphäre. Senckenberg 2018, pp. 22-25.

Bocherens, H., Pacher, M., 2019. Chapter 14. Isotopic analysis of cave bear collagen in Medvedia Cave (Slovakia). In (M. Sabol & G. Rabeder, Eds.): Medvedia Jaskyna Cave in the Slovensky raj Mts. Paleontological research 2007-2009. pp. 220-227.

* Bons, P.D., Bauer, C.C., Bocherens, H., de Riese, T., Drucker, D.G., Francken, M., Menéndez, L., Uhl, A., van Milligen, B.P., Wißing, C. 2019. Out of Africa by spontaneous migration waves. PLoS ONE 14(4): e0201998. journal.pone.0201998

* Crazzolara, C., Ebner, M., Platis, A., Miranda, T., Bange, J., Junginger, A. 2019. A new multicopter based unmanned aerial system for pollen and spores collection in the atmospheric boundary layer. Atmospheric Measurement Techniques 12, 1581-1598. 

* Díaz-Zorita Bonilla, M., Aranda Jiménez, G., Bocherens, H., Escudero Carillo, J., Sánchez Romero, M., Lozano Medina, Á, Alarcón García, E., Milesi García, L. In press. Multi-isotopic diet determination of Southeastern Iberian Megalithic populations: The cemeteries of El Barranquete and Panoría. Archaeological and Anthropological Science 11(8): 3681-3698. doi: 10.1007/s12520-018-0769-5.

* Etu-Sihvola, H., Bocherens, H., Drucker, D.G., Junno, A.P., Mannermaa, K., Oinonen, M., Uusitaol, J., Arppe, L., 2019. The dIANA database – isotopic baseline for paleodietary research in the Baltic Sea area. Journal of Archaeological Science: Reports 24: 1003-1013. doi: 10.1016/j.jasrep.2019.03.005. access to database

* Fuentes, R., Ono, R., Nakajima, N., Nishizawa, H., Siswanto, J., Aziz, N., Sriwigati, Sofian, H.O., Miranda, T., Pawlik, A. 2019. Technological and behavioural complexity in expedient industries: The importance of use-wear analysis for understanding flake assemblages. Journal of Archaeological Science, ISSN: 0305-4403, Vol: 112, Page: 105031.

Germonpré, M., Bocherens, H., Lázničková-Galetova, M., Sablin, M.V., 2019. La relation de l’homme au loup pendant le Paléolithique. Contes & Légendes 13: 38-41.

* Gretzinger, J., Molak, M., Reiter, E., Pfrengle, S., Urban, C., Neukamm, J., Blant, M., Conard, N.J., Cupillard, C., Dimitrijevic, V., Drucker, D.G., Hofman-Kaminska, E., Kowalczyk, R., Paunovic, M., Krajcarz, M.T., Krajcarz, M., Münzel, S.C., Peresani, M., Romandini, M., Rufí, I., Soler, J., Terlato, G., Krause, J., Bocherens, H., Schuenemann, V.J., 2019. Large-scale mitogenomic analysis of the phylogeographiy of the late Pleistocene cave bear. Scientific Reports 9: 10700 media report 1 media report 2 media report 3 media report 4

* Henry-Gambier, D., Rocher, P., Drucker, D., 2019. Étude et description de la sépulture de l’enfant de l’abri Labattut (vallon de Castelmerle, Sergeac, Dordogne, France). PALEO. Revue d'archéologie préhistorique, (30-1): 170-203. 

* Hofman-Kaminska, E., Bocherens, H., Drucker, D.G., Fyfe, R.M., Guminski, W., Makowiecki, D., Pacher, M., Piliciauskiene, G., Samojlik, T., Woodbridge, J., Kowalczyk, R., 2019. Adapt or die - response of large herbivores to environmental changes in Europe during the Holocene. Global Change Biology 25(9): 2915-2930. media report 1 media report 2

* Krajcarz, M.T., Krajcarz, M., Drucker, D.G., Bocherens, H., 2019.  Prey-to-Fox enrichment of 34S in bone collagen: Implications for palaeoecological studies. Rapid Communications in Mass Spectrometry 33: 1311-1317.

* Li D-Q, Zhou C-F, Li L, Yang J-T, Li L, Rabi M. 2019. The sinemydid  turtle Ordosemys from the Lower Cretaceous Mengyin Formation of  Shandong, China and its implication for the age of the Luohandong  Formation of the Ordos Basin. PeerJ 7:e6229

Ma, J., Wang, Y., Jin, C., Hu, Y., Bocherens, H., 2019. Ecological flexibility and differential survival of Pleistocene Stegodon orientalis and Elephas maximus in mainland southeast Asia revealed by stable isotopes (C, O) analysis. Quaternary Science Reviews 212: 33-44. j.quascirev.2019.03.021 media report 1 media report 2 (in Chinese)

* Massonne, T., Vasilyan, D., Rabi, M., Böhme, M., 2019. A new alligatoroid from the Eocene of Vietnam highlights an extinct Asian clade independent from extant Alligator sinensis. PeerJ 7e:e7562 

Prat, S., Péan, S., Crépin, L., Puaud, S., Verna, C., Drucker, D., Lázničková-Galetová, M., Patou-Mathis, M., Valladas, H., Der Plicht, J.V. and Yanevich, A., 2019, January. Les premiers Hommes anatomiquement modernes du sud-est de l’Europe: apport du site de Buran-Kaya III (Crimée). In Journée de la Société d'Anthropologie de Paris, session Cro-Magnon to the present day: peopling, population dynamics, human-environment interactions..

* Rey-Iglesia, A., García-Vazquez, A., Treadaway, E.C., van der Plicht, J., Baryshnikov, G.F., Szpak, P., Bocherens, H., Boeskorov, G.G., Lorenzen, E.D., 2019. Evolutionary history and palaeoecology of brown bear in North-East Siberia re-examined using ancient DNA and stable isotopes from skeletal remains. Scientific Reports 9:4462.

* Smith, O., Dunshea, G.J., Sinding, M.H.S., Fedorov, S., Germonpré, M., Bocherens, H., Gilbert, M.T.P., 2019. Shotgun sequencing of ancient RNA from late Pleistocene and historical canids shows tissue-specific transcriptome survival. PLOS Biology 17(7): e3000166. media report 1, media report 2

* Strani, F., Pushkina, D., Bocherens, H., Belluci, L., Sardella, R., DeMiguel, D., 2019.  Dietary adaptations of Early and Middle Pleistocene fossil equids from the Agnani basin (Frosinone, Central Italy). Frontiers in Ecology and Evolution vol.7, article 176. https:// doi: 10.3389/fevo.2019.00176

* Suraprasit, K., Jongautchariyakul, S., Yamee, C., Pothichaiya, C., Bocherens, H., 2019. New fossil and isotope evidence for the Pleistocene zoogeographic transition and hypothesized savanna corridor in Peninsular Thailand. Quaternary Science Reviews 221: 105861 media report 1 media report 2

* Terlato, G., Bocherens, H., Romandini, M., Nannini, N., Hobson, K.A., Peresani, M., 2019. Chronological and isotopic data support a revision for the timing of cave bear extinction in Mediterranean Europe. Historical Biology 31: 474-484. doi: 10.1080/08912963.2018.1448395 media report 1, media report 2

* Wißing, C., Rougier, H., Baumann, C., Comeyne, A., Crevecoeur, I., Drucker, D.G., Gaudzinski-Windheuser, S., Germonpré, M., Gómez-Olivencia, A., Krause, J., Matthies, T., Naito, Y.I., Posth, C., Semal, P., Street, M., Bocherens, H., 2019. Stable isotopes reveal patterns of diet and mobility in last Neandertals and first modern humans in Europe. Scientific Reports 9:4433 media report 1, media report 2


* Aranda Jiménez, G., Lozano Medina, Á., Sánchez Romero, M., Diaz-Zorita Bonilla, M., Bocherens, H., 2018. The chronology of the Megalithic funerary practices in South-Eastern Iberia: The necropolis of Panoria (Granada, Spain). Radiocarbon 60: 1-19. DOI: 10.1017/RDC.2017.96

* Bauer, C.C., Benazzi, S., Darlas, A., Harvati, K., 2018. Geometric morphometric analysis and internal structure measurements of the Neanderthal lower fourth premolars from Kalamakia, Greece. Quaternary International 497: 14-21. doi: 10.1016/j.quaint.2018.01.035

* Beck, J., Díaz-Zorita Bonilla, M., Bocherens, H., Diaz-del-Rio, P., 2018. Feeding a third millennium BC mega-site: Bioarchaeological analyses of palaeodiet and dental disease at Marroquíes (Jaén, Spain). Journal of Anthropological Archaeology 52: 23-43 doi: 10.1016/jaa.2018.07.001

* Bocherens, H., 2018. The rise of the anthroposphere since 50,000 years: an ecological replacement of megaherbivores by humans in terrestrial ecoystems? Frontiers in Ecology and Evolution 6:3 doi: 10.3389/fevo.2018.00003

* Bocherens, H., Cotte, M., Bonini, R., Straccia, P., Scian, D., Soibelzon, L., Prevosti, F.,J., 2018. Reply to “Comment on “Isotopic insight on paleodiet of ... by Bocherens et al. (Gondwana Research 48: 7-14).” Gondwana Research 58: 243-245. doi: 10.1016/

Bocherens, H., Drucker, D.G., Rivals, F. 2018. L’alimentation et l’environnement (micro-usure et isotopes) des mammouths et éléphants pléistocènes. In: Catalogue de l’exposition “Mémoires de Mammouths”, Musée national de Préhistoire Les Eyzies-de-Tayac, pp. 35-39.

* Chala-Aldana, D., Bocherens, H., Miller, C., Moore, K., Hodgins, G., Rademaker, K., 2018. Mobility and highland occupation strategies during the Early Holocene at the Cuncaicha rock shelter through strontium and oxygen isotopes. Journal of Archaeological Science: Reports 19: 811-827. doi:10.1016/j.jasrep.2017.10.023

Cintas-Peña, M., Sanjuán, L.G., Díaz-Zorita Bonilla, M., Herrero Corral, A.M., Robles Carrasco, S., 2018. La polación no adulta del asentamiento de la Edad del Cobre de Valencina de la Concepción (Sevilla): una aproximación demográfia, contextual y sociológia. Trabajos de Prehistoria 75(1): 85-108. doi: 10.3989/tp.2018.12205

* Cotte, M., Prevosti, F.,J., Hajdas, I., Scian, D., Straccia, P., Soibelzon, L., Bocherens, H., 2018. First AMS radiocarbon direct dates on bones from extinct megafauna in Camet Norte (Santa Clara del Mar, Buenos Aires Province, Argentina). Ameghiniana 55(1): 126-133. doi: 10.5710/AMGH.31.03.2017.3072

* Díaz-Zorita Bonilla, M., Beck, J., Bocherens, H., Díaz-del-Río, P., 2018. Isotopic evidence for human mobility at the Copper Age mega-site of Marroquíes Bajos (Jaén, Spain). Antiquity 92: 991-1007. doi: 10.15184/aqy.2018.33

* Drucker, D.G., Stevens, R.E., Germonpré, M., Sablin, M.V., Péan, S., Bocherens, H.,2018. Collagen stable isotopes provide insights into the end of the mammoth steppe in central East European plains during the Epigravettian. Quaternary Research 90: 457-469. doi: 10.1017/qua.2018.40media report 1, media report 2 

Drucker, D.G., Furtwängler, A., Schünemann, V., Huber, R., Reinhard, J., 2018. Durchleuchtet und analysiert. Ein Update zur Genetik, Isotopie und Radiografie des "letzten Zuger Mammuts". Tugium 34: 123-131. 

* Drucker, D.G., Valentin, F., Thevenet, C., Mordant, D., Cottiaux, R., Delsate, D., van Neer, W., 2018. Aquatic resources in human diet in the Late Mesolithic in Northern France and Luxembourg: insights from carbon, nitrogen and sulphur isotope ratios. Archaeol. Anthropol. Sci. 10: 351-368. doi: 10.1007/s12520-016-0356-6

Germonpré, M., Láznickova-Galetova, M., Sablin, M., Bocherens, H., 2018. Self-domestication or human control? The Upper Palaeolithic domestication of the dog. In: (J.-D. Vigne and C. Stépanoff, eds), Hybrid communities, biosocial approaches to domestiction and other trans-species relationships Routledge, pp. 39-64.

* Hofman-Kaminska, E., Bocherens, H., Borowik, T., Drucker, D.G., Kowalczyk, R., 2018. Stable isotope signatures of large herbivore foraging habits across Europe. PLoS ONE 13(1): e0190723.

* Hofman-Kaminska, E., Merceron, G., Bocherens, H., Makowiecki, D., Piliciauskiene, G., Ramdarshan, A., Berlioz, E., Kowalczyk, R., 2018. Foraging habitats and niche partitioning of European large herbivores during the Holocene. Palaeogeography, Palaeoclimatology, Palaeoecology 506: 183-195. doi: 10.1016/j.palaeo.2018.05.050

* Jiménez, G.A., Medina, À.L., Díaz-Zorita Bonilla, M., Romero, M.S., Carrillo, J.E., 2018. Cultural continuity and social resistance: The chronology of Megalithic Funerary Practices in Southern Iberia. European Journal of Achaeology 21(2): 192-216. doi: 10.1017/eaa.2017.42

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* Naito, Y.I., Chikaraishi, Y., Drucker, D.G., Ohkouchi, N., Semal, P., Wißing, C., Bocherens, H., 2018. Reply to “Comment on “Ecological niche of Neanderthals from Spy Cave revealed by nitrogen isotopes of individual amino acids in collagen.” (Journal of Human Evolution 93: 82-90.). Journal of Human Evolution 117: 56-60. doi: 10.1016/j.jhevol.2017.05.008

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Prat, S., Péan, S., Crépin, L., Puaud, S., Drucker, D.G., Lázničková-Galetová, M., Van der Plicht, J., Valladas, H., Verna, C., Patou-Mathis, M., Lebon, M., Yanevich, A., 2018. The first Anatomically Modern Humans from South-Eastern Europe. Contributions from the Buran-Kaya III site (Crimea). Bulletin de la Société d'Anthropologie de Paris 30: 169-179. doi: 10.3166/bmsap-2018-0032

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* Suraprasit, K., Bocherens, H., Chaimanee, Y., Somsak, P., Jaeger, J.-J., 2018. Middle Pleistocene glacial ecology and climate in Northeastern Thailand inferred from the stable isotope analysis of Khok Sung ungulate tooth enamel and the land mammal cenogram. Quaternary Science Reviews 193 : 24-42. doi: 10.1016/j.quascirev.2018.06.004

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* Dantas, M., Cherkinsky, A., Bocherens, H., Drefahl, M., Bernardes, C., França, L.M., 2017. Isotopic palaeoecology of the Pleistocene megamammals from the Brazilian Intertropical Region: Feeding ecology (δ13C, δ18O), niche breadth and overlap. Quaternary Science Reviews 170: 152-163. doi: 10.1016/j.quascirev.2017.06.030

Díaz-Zorita Bonilla, M., Roberts, C.A., Sanjuán, L.G., Pérez, V.H. 2017. Tomb 3 at La Pijotilla (Solana de los Barros, Badajoz, Spain): A bioarchaeological study of a Copper Age collective burial. In: (Tomé, T., Díaz-Zorita Bonilla, M., Silva A.M., Cunha, C., Boaventura, R., eds) Current approaches to collective burials in the Late European Prehistory. Archaeopress Archaeology, pp. 1-10.

Díaz-Zorita Bonilla, M., Knudson, K.J., Escudero Carrillo, J., Bocherens, H., García Sanjuán, L., 2017. Mobility patterns and paleodietary insights into humans and cattle at the copper age mega-site of Valencina (Seville, Spain) through δ18O and δ13C isotope analyses. Menga, Revista de Prehistoria de Andalucia 8: 52-68.

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* Trautmann, B., Diaz-Zorita Bonilla, M., Wißing, C., Bis-Woch, C., Bocherens, H., 2017. Reconstruction of socioeconomic status in the Medieval population of Grevenmacher (Luxembourg) based on growth, development and diet. International Journal of Osteoarchaeology 27: 947-957. doi: 10.1002/oa.2606


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Drucker, D.G., Rivals, F., Münzel, S.C., Bocherens, H., 2016. Stable isotope and microwear investigation on the mammoth (Mammuthus primigenius) of Kraków Spadzista: insights into diet and environment. In: (Wojtal, P., Wilczynski, J., Haynes, G., eds) A Gravettian site in southern Poland: Kraków Spadzista, pp. 189-202.

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Jiménez, G.A., Medina, À.L., Escudero Carrillo, J., Sánchez Romero, M., Alarcón García, E., Fernández Martín, S., Díaz-Zorita Bonilla, M., Barba Colmenero, V., 2016. Cronología y temporalidad de los recintos de fosos prehistóricos: el caso de Marroquíes Bajos (Jaén). Trabajos de Prehistoria 71(2): 231-250.

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Koehler, H., Wegmüller, F., Detrey, J., Diemer, S., Hauck, T., Pümpin, C., Sévèque, N., Stoetzel, E., Wuscher, P., Auguste, P., Bocherens, H., Lutz, M., Preusser, F., 2016. Fouilles de plusieurs occupations du Paléolithique moyen à Mutzig Rain (Alsace): Premiers résultats. Bulletin de la Société Préhistorique française 113: 429-474.

* Krajcarz, M., Pacher, M., Krajcarz, M.T., Laughlan, L., Rabeder, G., Sabol, M., Wojtal, P., Bocherens, H., 2016. Reconstruction of cave bear ecology based on geographical variation of stable isotopes in bone collagen during MIS 3 from Western to Eastern Europe. Quaternary Science Reviews 131: 51-72. doi: 10.1016/j.quascirev.2015.10.028 pdf

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* Polet, C., Bocherens, H., 2016. New insights into the marine contribution to ancient Easter Islanders’ diet. Journal of Archaeological Science: Reports 6: 709-719. doi: 10.1016/j.jasrep.2015.09.013

* Posth, C., Renaud, G., Mittnik, A., Drucker, D.G., Rougier, H., Cupillard, C., Valentin, F., Thevenet, C., Furtwängler, A., Wißing, C., Franken, M., Malina, M., Bolus, M., Lari, M., Gigli, E., Capecchi, G., Crevecoeur, I., Beauval, Flas, D., Germonpré, M., van der Plicht, J., Cottiaux, R., Gély, B., Ronchitelli, A., Wehrberger, K., Grigourescu, D., Svoboda, J., Semal, Caramelli, D., Bocherens, H., Harvati, K., Conard, N.J., Haak, W., Powell, A., Krause, J., in press. Pleistocene human mitochondrial genomes suggest late dispersal of non-Africans and a Late Glacial population turnover in Europe. Current Biology 26: 827-833.. doi: 10.1016/j.cub.2016.01.037 media report

Putelat, O., Logel. T., Bocherens, H., Geigl, E.-M., Guimaraes, S.L., Metz, B., 2016. Une chasse aristocratique dans le Ried Centre-Alsace au premier Moyen Âge: l’apport de l’archéozoologie à la connaissance du site d’Ostheim « Birgelsgaerten » (France, Haut-Rhin). In Peytremann (E.) ed. - Des Fleuves et des Hommes à l'époque mérovingienne. Territoire fluvial et société au premier Moyen Âge (5e-12e siècle). Actes des 33e Journées internationales d’Archéologie mérovingienne. Strasbourg, 28-30 septembre 2012, 42e suppl. à la RAE, pp 253-268.

* Quiles, A., Valladas, H., Bocherens, H., Delqué-Količ, E., Kaltnecker, E., van der Plicht, J., Delannoy, J.-J., Feruglio, V., Fritz, C., Monney, J., Philippe, M., Tosello, G., Clottes, J., Geneste, J.-M., 2016. A high-precision chronological model for the decorated Upper Palaeolithic cave of Chauvet-Pont d’Arc Cave, Ardèche, France. PNAS 113(7): 4670-4675. doi: 10.1073/pnas.1523158113 media report 1 media report 2

* Rawlence, N.J., Wood, J.R., Bocherens, H., Rogers, K.M., 2016. Dietary interpretations for extinct megafauna using coprolites, intestinal content and stable isotopes: complimentary or contradictory? Quaternary Science Reviews 142: 173-178. 10.1016/j.quascirev.2016.05.017

* Rougier, H., Crevecoeur, I., Beauval, C., Posth, C., Flas, D., Wißing, C., Furtwängler, A., Germonpré, M., Gómez-Olivencia, A., Semal, P., van der Plicht, J., Bocherens, H., Krause, J., 2016. Neandertal cannibalism at the “Troisième caverne” of Goyet (Belgium). Scientific Reports 6: 29005. 10.1038/srep29005

Schnitzler, A., Arbogast, R.-M., Bocherens, H., Drucker, D., Duchiron, M.-S., Granado, J., Putelat, O., Salzburger, W., Eberhard, A., Schmutz, A., Schlumbaum, A., 2016. Approche écologique et historique du cerf dans le Massif Vosgien. ISBN 978-2-85712-075-9, 56p.

* Wißing, C., Rougier, H., Crevecoeur, I., Germonpré, M., Naito Y.I., Semal, P., Bocherens, H., 2016. Isotopic evidence for dietary ecology of Neandertals in North-Western Europe. Quaternary International 411: 327-345 10.1016/j.quaint.2015.09.091 media report

* Young, M.T., Rabi, M., Bell, M.A., Foffa, D., Steel, S., Sachs, S., Peyer, K., 2016. Big-headed marine crocodyliforms and why we must be cautious when using extant species as body light proxies for long-extinct relatives. Paleontological Electronica 19.03.30A: 1-14.


* Bocherens, H., 2015. Isotopic tracking of large carnivore palaeoecology in the mammoth steppe. Quaternary Science Reviews 117: 42-71. doi: 10.1016/j.quascirev.2015.03.018

* Bocherens, H., Drucker, D.G., Germonpré, M., Lázničková-Galetová, M., Naito, Y., Wissing, C., Brůžek, J., Oliva, M., 2015. Reconstruction of the Gravettian food-web at Předmostí I using isotopic tracking of bone collagen. Quaternary International 359-360: 211-228. 10.1016/j.quaint.2014.09.044 media report 1, media report 2

* Bocherens, H., Hofman-Kamińska, E., Drucker, D.G., Schmölcke, U., Kowalczyk, R. 2015. European Bison as a Refugee Species? Evidence from Isotopic Data on Early Holocene Bison and Other Large Herbivores in Northern Europe. PLoS ONE 10(2): e0115090. doi:10.1371/journal.pone.0115090 media report 1, media report 2

* Brinkman, D.B., Densmore, M., Rabi, M., Ryan, M.J., & Evans, D.C. 2015. Marine turtles from the Late Cretaceous of Alberta, Canada. Canadian Journal of Earth Sciences. 52(8): 581-589. doi: 10.1139/cjes-2014-0189.

* Cupillard, C., Magny, M., Bocherens, H., Bridault, A., Bégeot, C., Bichet, V., Bossuet, G., Drucker, D., Gauthier, E., Jouannic, G., Millet, L., Richard, H., Rius, D., Ruffaldi, P., Walter-Simonnet, A.-V., 2015. Ecosystem and human responses to palaeoclimate changes in the Jura Mountains between 40 and 8 ka cal BP. Quaternary International 378: 40-72 doi: 10.1016/j.jhevol.2014.05.032 pdf

* Drucker, D.G., Vercoutère, C., Chiotti, L., Nespoulet, R., Crépin, L., Conard, N.J., Münzel, S.C., Higham, T., van der Plicht, J., Lázničková-Galetová, M., Bocherens, H., 2015. Tracking possible decline of woolly mammoth during the Gravettian in the Dordogne and the Swabian Jura using multi-isotope tracking (13C, 14C, 15N, 34S, 18O). Quaternary International 359-360: 304-317. link media report 1, media report 2

* Dyke, G., Vremir, M., Brusatte, S.L., Bever, G.S., Buffetaut, E., Chapman, S., Csiki-Sava, Z., Kellner, A., Martin, E., Naish, D., Norell, M., Osi, A., Pinheiro, F.L., Prondvai, E., Rabi, M., Rodrigues, T., Steel, L., Tong, H., Vila Nova, B.C., Witton, M., 2015. Thalassadromeus sebesensis - A new name for an old turtle. Comment on "Thalassodromeus sebesensis, an out of place and out of time Gondwanan tapejarid pterosaur", Grellet-Tinner and Codrea. Gondwana Research 27: 1680-1682. doi: 10.1016/

* Fontanals-Coll, M., Eulàlia Subirà, M., Díaz-Zorita Bonilla, M., Duboscq, S., Gibaja, J.F., 2015. Investigating palaeodietary and social differences between two differentiated sectors of a Neolithic community, La Bòbila Madurell-Can Gambús (north-east Iberian Peninsula). Journal of Archaeological Science: Reports 3: 160-170. do: 10.1016/jasrep.2015.06.013

* Immel, A., Drucker, D.G., Bonazzi, M., Jahnke, T.K., Münzel, S.C.. Schuenemann, V.J., Herbig, A., Kind, C.-J., Krause, J., 2015. Mitochondrial genomes of giant deers suggest their late survival in Central Europe. Scientific Reports 5: 10853 doi:10.1038/srep10853

* Julien, M.-A., Rivals, F., Serangeli, J., Bocherens, H., Conard, N. J., 2015. A new approach for deciphering between single and multiple accumulation events through the use of intra-tooth isotopic variations: Application to the Middle Pleistocene bone bed of Schöningen 13 II-4. Journal of Human Evolution 89: 114-128. doi: 10.1016/j.jhevol.2015.02.012

* Kuitems, M., van der Plicht, J., Drucker, D.G., van Kolfschoten, T., Palstra, S.W.L., Bocherens, H., 2015. Palaeoecological implications of C and N stable isotopes of well-preserved bone collagen from Schöningen, Germany. Journal of Human Evolution 89: 105-113 doi: 10.1016/j.jhevol.2015.01.008

* Ösi, A., Rabi, M., Makádi, L., 2015. An enigmatic crocodyloform tooth from the bauxites of western Hungary suggests hidden mesoeucrocodylian diversity in the Early Cretaceous European archipelago. PeerJ. doi:10.7717/peerj.1160

* Rabi, M. & Sebők, N. 2015. A revised Eurogondwana model: Late Cretaceous Notosuchian crocodyliforms and other vertebrate taxa suggest the retention of episodic faunal links between Europe and Gondwana during most of the Cretaceous. Gondwana Research 28: 1197-1211. doi:10.1016/

* Rivals, F., Julien, M.-A., van Kolfschoten, T., Serangeli, J., Bocherens, H., Drucker, D.G., Conard, N. J., 2015. Investigation of equid paleodiet from Schöningen 13 II-4 through dental wear and isotopic analyses: Archaeological implications. Journal of Human Evolution 89: 129-137 doi: 10.1016/j.jhevol.2014.04.002 pdf

Schneider, M., Drucker, D.G., Buuveibaatar, B., Bocherens, H., 2015. Tracking saiga horns to the population of their origin with stable isotopes. Saiga News 19: 21-23.

* Wißing, C., Matzerath, S., Turner, E., Bocherens, H., 2015. Palaeoecological and climatic implications of stable isotope results from late Pleistocene bone collagen, Ziegeleigrube Coenen, Germany. Quaternary Research. 84: 96-105. doi:10.1016/j.yqres.2015.05.005

* Zhou, C.-F. & Rabi, M. 2015. A sinemydid turtle from the Jehol Biota provides insights into the basal divergence of crown-turtles. Scientific Reports 5, 16299.


* Aiglstorfer, M., Bocherens, H., Böhme, M., 2014. Large Mammal Ecology in the late Middle Miocene locality Gratkorn (Austria). Palaeobiodiversity and Palaeoenvironments 94: 189–213. pdf

Bocherens, H., 2014. Diet and ecology of the Scladina Neanderthal child: insights from stable isotopes. In: (M. Toussaint; D. Bonjean, Eds.) The Scladina I-4A Juvenile Neandertal. ERAUL 134: 351-362.

* Bocherens, H., Baryshnikov, G., van Neer, W., 2014. Were bears or lions involved in salmon accumulation in the Middle Palaeolithic of the Caucasus? An isotopic investigation in Kudaro 3 cave. Quaternary International 339-340: 112-118. DOI: 0.1016/j.quaint.2013.06.026 pdf media report 1, media report 2, media report 3, report in Jahresbericht Uni Tübingen 2013

* Bocherens, H., Bridault, A., Drucker, D.G., Hofreiter, M., Münzel, S.C., Stiller, M., van der Plicht, J., 2014. The last of its kind? Radiocarbon, ancient DNA and stable isotope evidence from a late cave bear from Rochedane (France). Quaternary International 339-340: 179-188. doi: 10.1016/j.quaint.2013.05.021 pdf

* Bocherens, H., Drucker, D.G., Madelaine, S., 2014. Evidence for a 15N positive excursion in terrestrial foodwebs at the middle to upper Palaeolithic transition in South-western France: implication for early modern human palaeodiet and palaeoenvironment. Journal of Human Evolution 69: 31-43. pdf media report

Bocherens, H., Drucker, D.G., Wissing, C., 2014. Die Mammutsteppe: Isotopenuntersuchungen in einem vergangenen Ökosystem. Natur - Forschung - Museum 144(7/8): 226-231. pdf

* Bocherens, H., Grandal-d'Anglade, A., Hobson, K.A., 2014. Pitfalls in comparing modern hair and fossil bone collagen C and N isotopic data to reconstruct ancient diets: a case study with cave bears (Ursus spelaeus). Isotopes in Health and Environmental Sciences 50(3): 291-299. DOI: 1080/10256016.2014.890193 pdf

* Corsini, J.A., Böhme, M., Joyce, W.G., 2014. Reappraisal of Testudo antiqua (Testudines, Testudinidae) from the Miocene of Hohenhöwen, Germany. Journal of Paleontology 88(5): 948-966.

* Drucker, D.G., Bocherens, H., Péan, S. 2014. Isotopes stables (13C, 15N) du collagène des mammouths de Mezhyrich: implications paléoécologiques. L’Anthropologie 118(5): 518-537. doi: 10.1016/j.anthro.2014.04.001 link

* França, L., Dantas, M., Bocchiglieri, A., Cherckinsky, A., Ribeiro, A., Bocherens, H., 2014. Chronology and ancient feeding ecology of two Upper Pleistocene megamammals from the Brazilian Intertropical Region. Quaternary Science Reviews 99: 78-83. pdf

*Mahboubi, S., Bocherens, H., Scheffler, M., Benammi, M., Jaeger, J.-J., 2014. Was the Early Eocene proboscidean Numidotherium koholense semi-aquatic or terrestrial? Evidence from stable isotopes and bone histology. Comptes Rendus Palevol 13: 501-509. doi:10.1016/j.crpv.2014.01.002 pdf

Maier W, Werneburg I (Eds.) Schlüsselereignisse der organismischen Makroevolution. Scidinge Hall Verlag Zürich, 411p link

* Martin, J.E, Rabi, M., Csiki, Z. & Vasile, S. 2014. Cranial morphology of Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae) and the widespread occurrence of Theriosuchus in the Late Cretaceous of europe. Journal of Paleontology 88(3): 444-456.

* Münzel, M.C., Rivals, f., Pacher, M., Döppes, D., Rabeder, G., Conard, N.J., Bocherens, H., 2014. Behavioural ecology of Late Pleistocene cave bears (Ursus spelaeus, U. ingressus): insights from stable isotopes (C, N, O) and tooth microwear. Quaternary International 339-340: 148-163. pdf

* Püntener, C., Billon-Bruyat, J.-P., Bocat, L., Berger, J.-P., Joyce, W.G. 2014. Taxonomy and phylogeny of the turtle Tropidemys langii Rütimeyer, 1873 based on new specimens from the Kimmeridgian of the Swiss Jura Mountains. Journal of Vertebrate Paleontology 34(2): 353-374. link

* Pushkina, D., Bocherens, H., Ziegler, R., 2014. Unexpected palaeoecological features of the middle and late Pleistocene large herbivores in south-western Germany revealed by stable isotopic abundances in tooth enamel. Quaternary International 339-340: 164-178. pdf

* Rabi, M. Sukhanov, V.B., Egorova, V., Danilov, I. & Joyce, W.G. 2014. Osteology, relationships, and ecology of Annemys (Testudines: Eucryptodira) from the Late Jurassic of Shar Teg, Mongolia and a phylogenetic definition Of Xinjiangchelyidae, Sinemydidae, and Macrobaenidae. Journal of Vertebrate Paleontology 34 (2): 327–352. doi: 10.1080/02724634.2013.807274

* Soibelzon, L.H., Grinspan, G.A., Bocherens, H., Acosta, W.G., Jones, W.W., Blanco, R.E., Prevosti, F.J., 2014. South American giant short-faced bear (Arctotherium angustidens) diet: evidence from pathology, morphology, stable isotopes and biomechanics. Journal of Palaeontology 88(6): 1240-1250. link

* Stiller, M., Molak, M., Prost, S., Rohland, N., Pacher, M., Rabeder, G., Baryshnikov, G., Rosendahl, W., S., Münzel, Bocherens, H., Grandal-d'Anglade, A., Hilpert, B., Germonpré, M., Stasyk, O., Pinhasi, R., Tintori, A., Ho, S. Y. W., Hofreiter, M., Knapp, M., in press. Mitochondrial DNA diversity and evolution of the Pleistocene cave bear complex. Quaternary International 339-340: 224-231. pdf

* Suraprasit, K., Chaimanee, Y., Bocherens, H., Chavasseau, O., Jaeger, J.-J., 2014. Middle Miocene Cervidae (systematics and phylogeny) and paleoenvironmental implications of Mae Moh basin by using the stable isotope analysis of the herbivore tooth enamel. Journal of Vertebrate Paleontology 34(1): 179-194. doi: 10.1080/02724634.2013.789038

* Tsutaya, T., Naito, Y.I., Ishida, H., Yoneda, M., 2014. Carbon and nitrogen isotope analyses of human and dog diet in the Okhotsk culture: perspectives from the Moyoro site, Japan. Anthropological Science 122(2): 89-99 doi: 10.1537/ase.140604 pdf

* Vercoutère, C., Crépin, L., Drucker, D.G., Chiotti, L., Henry-Gambier, D., Nespoulet, R., 2014. Deer (Rangifer tarandus and Cervus elaphus) remains from the final Gravettian of the abri Pataud and their importance to humans. In: Deer and People: Past, Present and Future, K. Baker, R. Carden, R. Madgwick (eds.), Oxford, Oxbow Books, Windgather Press, pp. 145-158.

Werneburg I. 2014. Konvergente Evolution herbivorer Landwirbeltiere [Review]. In: Maier W, Werneburg I (Eds.) Schlüsselereignisse der organismischen Makroevolution. Scidinge Hall Verlag Zürich, pp. 295-331.

Werneburg I. 2014. Vom Ursprung des Menschen bis zur Neolithischen Revolution [Review]. In: Maier W, Werneburg I (Eds.) Schlüsselereignisse der organismischen Makroevolution. Scidinge Hall Verlag Zürich, pp. 361-411.

* Zhou, C.-F., Rabi, M. & Joyce, W.G. 2014. A new specimen of Manchurochelys manchoukuoensis from the Early Cretaceous Jehol Biota of Chifeng, Inner Mongolia, China and the phylogeny of Cretaceous basal eucryptodiran turtles. BMC Evolutionary Biology. 14 (77): 1–16.


* Bibi, F., Souron, A., Bocherens, H., Uno, K., Boisserie, J.R., in press. Ecological Change in the Lower Omo Valley around 2.8 Ma. Biology Letters 9(1): 20120890. doi: 10.1098/rsbl.2012.0890 pdf review in Nature

* Bocherens, H. & Drucker, D.G. Stable isotopes in terrestrial teeth and bones. in: (S. Elias, Ed.) Encyclopedia of Quaternary Sciences, Elsevier, 2nd Edition, vol. 1, pp. 304-314, Amsterdam: Elsevier. pdf

Bocherens, H., Germonpré, M., Toussaint, M., Semal, P. 2013. XVII. Stable isotopes. In: (P. Semal & A. Hauzeur, Eds), Spy Cave: State of 120 Years of Pluridisciplinary Research on the Betche-aux-Rotches from Spy, pp. 357-371.

Carabajal AP, Sterli J, Müller J, Joyce WG, Werneburg I (2013). Endocranial reconstructions of extinct turtles using μ-CT scans: new insights into brain and inner ear anatomy. 10th International Congress of Vertebrate Morphology (ICVM), in Barcelona / Spain, 8. – 12. July 2013, Anatomical Record 295 (Special Feature): page 207.

*Delfino, M., Scheyer, T. M., Chesi, F., Fletcher, T., Gemel, R., Rabi, M. & Salisbury, S. W. 2013. Gross morphology and microstructure of type locality ossicles of Psephophorus polygonus Meyer, 1847 (Testudines, Dermochelyidae). Geological Magazine 150 (5): 767–782. pdf

Díaz-Zorita Bonilla, M. 2013. Bioarqueología de las prácticas funerarias del yacimiento de la Edad del Cobre de Valencina de la Concepción-Castilleja de Guzmán: revisión de las investigaciones (State of the art of the bioarchaeology of the funerary practices at the archaeological site of Valencina de la Concepción). In García Sanjuán, L., Hurtado Pérez, V., Vargas Jiménez, J. M., Ruiz Moreno, T. and Cruz-Auñón Briones, R. (Editores): Valencina Prehistórica. Actas del Congreso Conmemorativo del Descubrimiento de La Pastora (1860-2010). Sevilla, Universidad de Sevilla.

Drucker, D.G., Bocherens, H., 2013. Reconstitution de l’alimentation des coptes d’Antinoe par les analyses d’isotopes stables du carbone, de l’azote et du soufre de la kératine de cheveux de momies. In: (Y. Lintz & M. Coudert, eds) Collections Egyptiennes d’Antinoé (momies, tissues, céramiques et autres antiques). Envois de l’Etat et dépôts du musee du Louvre (1895-1925), p. 82-85. pdf

* Ecker, M., Bocherens, H., Julien, M.-A., Rivals, F., Raynal, J.-P., Moncel, M.-H., 2013. Middle Pleistocene ecology and Neanderthal subsistence: Insights from stable isotope analyses in Payre (Ardèche, France). Journal of Human Evolution 65: 363-373. pdf

García Sanjuán, L. and Díaz-Zorita Bonilla, M. 2013. Prácticas funerarias en estructuras negativas del asentamiento prehistórico de Valencina de la Concepción (Sevilla, España): análisis contextual y osteoarqueológico (Analysis of the osteoarchaeological remains documented at negative structures at the Copper Age site of Valencina de la Concepción). In García Sanjuán, L., Hurtado Pérez, V., Vargas Jiménez, J. M., Ruiz Moreno, T. and Cruz-Auñón Briones, R. (Editors):Valencina Prehistórica. Actas del Congreso Conmemorativo del Descubrimiento de La Pastora (1860-2010). Sevilla, Universidad de Sevilla.

* Joyce, J.C., Werneburg, I., Lyson, T.R., 2013. The hooked element in the pes of turtles (Testudines): a global approach to exploring primary and secondary homology. Journal of Anatomy 223: 421-441. pdf

* Joyce, W.G., Parham, J.F., Lyson, T.R., Warnock, R.C.M., Donoghue, P.C.J. 2013. A divergence dating analysis of turtles using fossil calibrations: an example of best practice. Journal of Paleontology 87(4): 612-634. link

Leduc, C., Bridault, A., Souffi, B., David, E., Drucker, D.G., 2013. Apports et limites de l’étude des vestiges fauniques à la caractérisation d’un site mésolithique de plein air à Paris : « 62 rue Henry-Farman » (15e arrondissement ; Ile-de-France). Bulletin de la Société Préhistorique française 110(2), 257-280.

* Lyson, T.R., Bhullar, B.-A.S., Bever, G.S., Joyce, W.G., de Queiroz, K., Abzhanov, A., Gauthier, J.A. 2013. Homology of the enigmatic nuchal bone reveals novel reorganzation of the shoulder girdle in the evolution of the turtle shell. Evolution and Development 15(5): 317-325.

Matzerath, S., Turner, E., Fischer, P., van der Plicht, J., Bocherens, H., Wißing, C., Kels, K., 2013. Interdisziplinäre Forschungen zu eiszeitlichen Großsäugern aus der Ziegeleigrube Coenen. Archäologie im Rheinland 2012, Theiss, pp. 67-69. pdf

* Naito, Y., Chikaraishi, Y., Ohkouchi, N., Yoneda, M., 2013. Evaluation of carnivory in inland Jomon hunter-gatherers based on nitrogen isotopic compositions of individual amino acids in bone collagen. Journal of Archaeological Science 40(7): 2913-2923.

* Naito, Y.I., Chikaraishi, Y., Ohkouchi, N., Drucker, D.G., Bocherens, H. 2013. Nitrogen isotopic composition of collagen amino acids as an indicator of aquatic resource consumption: insights from Mesolithic and Epipalaeolithic archaeological sites in France. World Archaeology 45: 338-359. DOI: 10.1080/00438243.2013.820650 pdf

* Péan, S., Puaud, S., Crépin, L., Prat, S., Quiles, A., van der Plicht, J., Valladas, H., Stuart, A.J., Drucker, D.G., Patou-Mathis, M., Lanoe, F., Yanevich, A., 2013. The Middle to Upper Paleolithic sequence of Buran-Kaya III (Crimea, Ukraine): New stratigraphic, paleoenvironmental, and chronological results. Radiocarbon, 55: 1454-1469. pdf

* Polachowski KM, Werneburg I (2013). Late embryos and bony skull development in Bothropoides jararaca (Serpentes, Viperidae). Zoology 116 (1): 36-63. doi: 10.1016/j.zool.2012.07.003.

* Rabi M, Zhou C-F, Wings O, Ge S & Joyce WG 2013. A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles. BMC Evolutionary Biology 13:203, 1-28. pdf

Robles Carrasco, S. and Díaz-Zorita Bonilla, M. 2013. Análisis bioarqueológico de tres contextos‐estructuras funerarias del sector PP4 – Montelirio del yacimiento de Valencina de la Concepción ‐ Castilleja de Guzmán (Sevilla) (Bioarchaeological analysis of three funerary contexts-structures at PP4-Montelirio sector from Valencina-Castilleja site).In García Sanjuán, L.; Hurtado Pérez, V.; Vargas Jiménez, J. M.; Ruiz Moreno, T. y Cruz-Auñón Briones, R. (Editores):Valencina Prehistórica. Actas del Congreso Conmemorativo del Descubrimiento de La Pastora (1860-2010). Sevilla, Universidad de Sevilla.

* Scheyer TM, Werneburg I, Mitgutsch C, Delfino M, Sánchez-Villagra MR (2013). Chapter 6: Three ways to tackle the turtle: integrating fossils, comparative embryology and mircoanatomy. In: Morphology and Evolution of Turtles. Vertebrate Paleobiology and Paleoanthropology Series. Editors: Gardner J, Brinkman D, Holroyd P. Springer Verlag, Dortrecht, Heidelberg, New York, London, pp. 63-70. pdf

Semal, P., Hauzeur, A., Rougier, H., Crevecoeur, I., Germonpré, M., Pirson, S., Haesaerts, P., Jungels, C., Flas, D., Toussaint, M., Maureille, B., Bocherens, H., Higham, T., Van Der Plicht, J., i2013. Radiocarbon dating. In: Rougier, H., Semal, P. (Eds), Spy Cave. State of 125 years of pluridisciplinary research on the Betche aux Rotches from Spy (Jemeppe-sur-Sambre, Province of Namur, Belgium). Royal Belgian Institute of Natural Sciences, pp. 331-356. pdf

* Vasilyan, D., Böhme, M., Chkhikvadze, V.M., Semenov, Y.A., Joyce, W.G. 2013. A new giant salamander (Urodela, Pancryptobrancha) from the Miocene of Eastern Europe (Grytsiv, Ukraine). Journal of Vertebrate Paleontology 33:301–318.

* Vercoutère, C., Guérin, C., Crépin, L., Richardin, P., Gandolfo, N., Vincent, J., Marsac, J., Cersoy, S., Rousselière, H., Walter, P., Brunelle, A., Nowik, W., Brissaud, D., Drucker, D.G., van der Plicht, J., Patou-Mathis. M., Vialet, A., 2013. Pluridisciplinary research on a woolly rhinoceros skeleton, Coelodonta antiquitatis (Blumenbach, 1799), from the Institut de paléontologie humaine (Paris, France). L’Anthropologie 117: 1-47. pdf

* Werneburg, I (2013). Jaw musculature during the dawn of turtle evolution. Organisms, Diversity & Evolution 13: 225-254. pdf

* Werneburg I (2013). The tendinous framework in the temporal skull region of turtles and considerations about its morphological implications in amniotes: a review. Invited Review to Zoological Science 31(3): 141-153.

* Werneburg I, Maier W, Joyce WG 2013. Embryonic remnants of intercentra and ribs in the neck of turtles. Biology Open 2: 1103-1107. doi: 10.1242/bio.20135439

* Werneburg I, Tzika AC, Hautier L, Asher RJ, Milinkovitch MC, Sánchez-Villagra MR (2013). Development and embryonic staging in non-model organisms: the case of an afrotherian mammal. Invited paper to the special issue "Vertebrate Evolutionary Developmental Biology" in The Journal of Anatomy 222:2-18. pdf

Werneburg I, Wilson LAB, Joyce WG (2013). Evolution of neck vertebral shape and neck retraction at the transition to modern turtles: a geometric morphometric approach. 10th International Congress of Vertebrate Morphology (ICVM), in Barcelona / Spain, contributed talk in the symposium 'Quantifying evolutionary development using non-model organisms: integrating metrical frameworks, gene expression, and morphology' (organized by Laura A.B. Wilson and Ingmar Werneburg), 8. – 12. July 2013, Anatomical Record 295 (Special Feature): page 135.

Werneburg I, Joyce W, Maier W (2013). Embryonic remnants of intercentra and ribs in a pleurodire turtle - a case of recapitulation and functional reduction. 10th International Congress of Vertebrate Morphology (ICVM), in Barcelona / Spain, 8. – 12. July 2013, Anatomical Record 295 (Special Feature): page 257.

Werneburg I, Hinz J, Gumpenberger M, Volpato V, Natchev N, Joyce WG (2013). Peculiarities of turtle body plan: the origin and evolution of neck retraction mechanisms. 10th International Congress of Vertebrate Morphology (ICVM), in Barcelona / Spain. Invited talk to the symposium 'Origin and Evolution of Turtles' (organized by Jacqueline E. Moustakas and Scott Gilbert), 8. – 12. July 2013, Anatomical Record 295 (Special Feature): page 173.

* Yeakel, J.D., Guimarães P.R. Jr., Bocherens, H., Koch, P.L., 2013. The impact of climate change on the structure of Pleistocene food webs across the mammoth steppe. Proceedings of the Royal Society B: Biological Sciences 280: 20130239. doi:10.1098/rspb.2013.0239. pdf media report 1 media report 2


* Drucker, D.G., Bridault, A., Cupillard, C., 2012. Environmental context of the Magdalenian settlement in the Jura Mountains using stable isotope tracking (13C, 15N, 34S) of bone collagen from reindeer (Rangifer tarandus). Quaternary International 272-273, 322-332. pdf

* Drucker, D.G., Hobson, K.A, Münzel, S.C., Pike-Tay, A., 2012. Intra-individual variation in stable carbon (δ13C) and nitrogen (δ15N) isotopes in mandibles of modern caribou of Qamanirjuaq (Rangifer tarandus groenlandicus) and Banks Island (Rangifer tarandus pearyi): implications for tracing seasonal and temporal changes in diet. International Journal of Osteoarchaeology 22: 494–504. pdf

* Jones MEH, Werneburg I, Curtis N, Penrose R, O'Higgins P, Fagan MJ, Evans SE. 2012. The head and neck anatomy of sea turtles (Cryptodira: Chelonioidea) and skull shape in Testudines. PLoS ONE 7(11): e47852. doi:10.1371/journal.pone.0047852. pdf

* Joyce, W.G., Micklich, N., Schaal, S.F.K., Scheyer, T.M. Caught in the act: the first record of copulating fossil vertebrates. Biology Letters 8:846–848. media report 1 media report 2

* Joyce, W.G., Petričević, A., Lyson, T.R., Czaplewski, N.J. 2012. A new box turtle from the Miocene/Pliocene boundary (Latest Hemphillian) of Oklahoma and a refined chronology of box turtle diversification. Journal of Paleontology 86:177–190.

* Joyce, W.G., Sterli, J. 2012. Congruence, non-homology, and the phylogeny of basal turtles. Acta Zoologica 93:149–159.

Joyce WG, Corsini JA, Werneburg I, Rabi M (editors) (2012). Symposium on Turtle Evolution 2012. Program and Abstracts. TOBIAS-lib Hochschulserver der Universität Tübingen, 52 pp.

* Julien, M.-A., Bocherens, H., Burke, A., Drucker, D.G., Patou-Mathis, M., Krotova, A., Péan, S., 2012. Were European steppe bison migratory? 18O, 13C and Sr intra-tooth isotopic variations applied to a paleoethological reconstruction. Quaternary International 271: 106-119. DOI: 10.1016/j.quaint.2012.06.011 pdf

*Lyson, T.R., Joyce, W.G. 2012. Evolution of the turtle bauplan: the topological relationship of the scapula relative to the ribcage. Biology Letters 8:1028–1031.

Natchev N, Werneburg I (2012). Considerations on the evolution of feeding systems in turtles. Symposium on Turtle Evolution, University of Tübingen, Germany, 2-5 June 2012. Abstract published in: W.G. Joyce, J.A. Corsini, I. Werneburg, M. Rabi (eds.). TOBIAS-lib Hochschulserver der Universität Tübingen, page 32.

Ősi, A., Rabi, M., Makádi, L., Szentesi, Z., Botfalvai, G. & Gulyás, P. 2012. The Late Cretaceous continental vertebrate fauna from Iharkút (Western Hungary, Central Europe): a review, pp. 533–570 In “Tribute to Charles Darwin and the Bernissart Iguanodons: new perspectives of vertebrate evolution and Early Cretaceous ecosystems” (ed. Pascal Godfroit), Indiana University Press, Bloomington.

Pacher, M., Bocherens, H., Döppes, D., Frischauf, C., Rabeder, G., 2012. First results of stable isotopes from Drachenloch and Wildenmannlisloch, Swiss Alps. Braunschweiger Naturkundliche Schriften 11: 101-110. pdf

* Parham, J.F., et al., Joyce, W.G., et al. 2012. Best practices for justifying fossil calibrations. Systematic Biology 61:346–359.

Rabi, M., Vremir, M. & Tong, H. 2012. Preliminary overview of Late Cretaceous turtle diversity in eastern Central Europe (Austria, Hungary, and Romania), pp. 307–336 In “Morphology and Evolution of Turtles: Origin and Early Diversification” (eds: D. B. Brinkman, P. A. Holroyd, and J. D. Gardner), Springer, Dordrecht.

Rabi, M., Tong, H. & Botfalvai, G. 2012. A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini. Geological Magazine 149: 662–674. pdf

Rougier, H., Crevecoeur, I., Beauval, C., Bocherens, H., Flas, D., Germonpré, M., Semal, P., van der Plicht, J. 2012. New data from an old site: Neandertals at Goyet (Belgium) and their mortuary behavior. American Journal of Physical Anthropology, Abstracts of the 2012 Conference. p. 252-253.

Rougier, H., Crevecoeur, I., Beauval, C., Bocherens, H., Flas, D., Germonpré, M., Semal, P., van der Plicht, J. 2012. Neandertals at the troisième caverne of Goyet (Belgium) and their mortuary behavior. Bull. Mém. Soc. Anthropol. Paris. 23: S33-S34.

Sánchez-Villagra, M.R. (2012). Embryonen aus der Tiefenzeit. Was Fossilien über die Evolution von Wachstumsprozessen und die Entwicklungsgenetik erzählen. German translation by I. Werneburg. vdf Hochschulverlag AG an der ETH Zürich. 239 pages. ISBN 3728134694. Original Titel: Sánchez, M.R. Embryos in Deep Time: The Rock Record of Animal Development in Evolution (Indiana Press 2012)

* Werneburg, I 2012. Temporal bone arrangements in turtles: an overview. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution 318: 235-249.

Werneburg I, Lyson TR, Joyce WG (2012). An integrative study on the anatomy and development of the ‘hooked fifth metatarsal’ in the turtle pes. Symposium on Turtle Evolution, University of Tübingen, Germany, 2-5 June 2012. Abstract published in: W.G. Joyce, J.A. Corsini, I. Werneburg, M. Rabi (eds.). TOBIAS-lib Hochschulserver der Universität Tübingen, page 48

*Wings, O., Rabi, M., Schneider, J.W., Schwermann, L., Sun, G., Zhou, C.-F., Joyce, W.G. 2012. An enormous Jurassic turtle bone bed from the Turpan Basin of Xinjiang, China. Naturwissenschaften 99:925–935. pdf media report


​​​​​​​* Bocherens, H. 2011. Diet and ecology of Neanderthals: Insights from bone and tooth biogeochemistry. in: (N.J. Conard & J. Richter, Eds.), Neanderthal lifeways, subsistence and technology. Vertebrate Paleobiology & Paleoanthropology (series editors Eric Delson and Ross MacPhee). Heidelberg, New York – Springer, pp. 73-85. pdf media report

* Bocherens, H., Drucker, D.G., Bonjean, D., Bridault, A., Conard, N.J., Cupillard, C., Germonpré, M., Höneisen, M., Münzel, S.C., Napierrala, H., Patou-Mathis, M., Stephan, E., Uerpmann, H.-P., Ziegler, R. 2011. Isotopic evidence for dietary ecology of cave lion (Panthera spelaea) in North-western Europe: prey choice, competition and implications for extinction. Quaternary International 245: 249-261. pdf media report

* Bocherens, H., Drucker, D.G., Taubald, H. 2011. Preservation of bone collagen sulphur isotopic compositions in an early Holocene river-bank archaeological site. Palaeogeography, Palaeoclimatology, Palaeoecology 310: 32-38. DOI: 10.1016/j.palaeo.2011.05.016. pdf

* Bocherens, H., Pacher, M. 2011. Late Quaternary mammal ecology: insight from new approaches. Quaternary International 245: 183–185. DOI: 10.1016/j.quaint.2011.07.030 pdf

* Bocherens, H., Sandrock, O., Kullmer, O., Schrenk, F. 2011. Hominin palaeoecology in Late Pliocene Malawi: Insights from isotopes (13C, 18O) in mammal teeth. South African Journal of Science 107(3/4): 96-101. DOI: 10.4102/sajs.v107i3/4.331 pdf

* Bocherens, H., Stiller, M., Hobson, K.A., Pacher, M., Rabeder, G., Burns, J.A., Tütken, T., Hofreiter, M. 2011. Niche partitioning between two sympatric genetically distinct cave bears (Ursus spelaeus and Ursus ingressus) and brown bear (Ursus arctos) from Austria: isotopic evidence from fossil bones. Quaternary International. 245: 238-248. DOI: 10.1016/j.quaint.2010.12.020 pdf

* Drucker, D.G., Bridault, A., Cupillard, C., Hujic, A., Bocherens, H., 2011. Evolution of habitat and environment of red deer (Cervus elaphus) during the Late-glacial and early Holocene in eastern France (French Jura and the western Alps) using multi-isotope analysis (δ13C, δ15N, δ18O, δ34S) of archaeological remains. Quaternary International 245: 268-278. DOI:10.1016/j.quaint.2011.07.019 pdf

* Drucker, D.G., Kind, C.-J., Stephan, E. 2011. Chronological and ethological information on the early Holocene reindeer in NW Europe using radiocarbon and stable isotope analysis: case study from southern Germany. Quaternary International. 245: 218-224. doi:10.1016/j.quaint.2011.05.007 pdf

* Drucker, D.G., Madelaine, S., Morala, A., 2011. Les derniers rennes de Dordogne : nouvelles données chronologiques et environnementales par l’étude isotopique du collagène (13C, 14C, 15N). Paléo 22, 85-100. pdf

* Gauthier, J.A., Nesbitt, S.J. Joyce, W.G., Schachner, E. A 2011. nearly complete skeleton of a bipedal stem crocodilian and innovation in archosaur locomotion. Bulletin of the Peabody Museum of Natural History 52:107–126.

* Jaeger, J.-J., Soe, A. N., Chavasseau, O., Coster, P., Emonet, E.-G., Guy, F., Lebrun, R., Maung, A., Khyaw, A. A., Shwe, H., Tun, S. T., Linn Oo, K., Rugbumrung, M. Bocherens, H., Chaivanich, K., Tafforeau, P., Chaimanee, Y. 2011. First hominoid from the Late Miocene of the Irrawaddy Formation (Myanmar). PLOS ONE 6(4): e17065. doi:10.1371/journal.pone.0017065 pdf

* Joyce, W.G., Chapman, S.D., Moody, R.T.J., Walker, C.A. 2011. The skull of the solemydid turtle Helochelydra nopscai from the Early Cretaceous (Barremian) of the Isle of Wight (UK) and a review of Solemydidae. Special Papers in Paleontology 86:75–97.

* Joyce, W.G., Lyson, T.R. 2011. New material of Gilmoremys lancensis nov. comb. (Testudines: Trionychidae) from the Hell Creek Formation and the diagnosis of plastomenid turtles. Journal of Paleontology 85:442-459.

* Knauss, G., Joyce, W.G., Lyson, T.R., Pearson, D. 2011. A new kinosternoid from the Late Cretaceous Hell Creek Formation of North Dakota and Montana and the origin of the Dermatemys mawii lineage. Paläontologische Zeitschrift 85:125–142.

Ksepka, D.T., et al., Joyce, W.G., et al. 2011. Synthesizing and databasing fossil calibrations: divergence dating and beyond. Biology Letters 7:801-803.

Lyson, T.R., Bercovici, A., Chester, S., Sargis, E., Pearson, D., Joyce, W.G. 2011. Dinosaur extinction: closing the “3 m gap.” Biology Letters 7:925-928. media report 1 media report 2

* Lyson, T.R., Joyce, W.G. 2011. Cranial anatomy and phylogenetic placement of the enigmatic turtle Compsemys victa. Journal of Paleontology 85:789–801.

* Lyson, T.R., Joyce, W.G., Knauss, G.E, Pearson, D.A. 2011. Boremys (Testudines: Baenidae) from the Latest Cretaceous and Early Paleocene of North Dakota: an 11 million year range extension and an additional K/T survivor. Journal of Vertebrate Paleontology 31:729–737.

* Münzel, S.C., Hofreiter, M., Stiller, M., Mittnik, A., Conard, N.J., Bocherens, H., 2011. Pleistocene bears in the Swabian Jura (Germany): genetic replacement, ecological displacement, extinctions and survival. Quaternary International 245: 225-237. DOI: 10.1016/j.quaint.2011.03.060 pdf

* Prat, S., Péan, S.C., Crépin, L., Drucker, D.G., Puaud, S.J., Valladas. H., Láznicková-Galetová, M., van der Plicht, J., Yanevitch, A., 2011. The oldest anatomically modern humans from Far Southeast Europe: Direct dating, culture and behavior. PLOS One 6(6): e20834. doi:10.1371/journal.pone.0020834. pdf media report 1 media report 2


Blant, M., Bocherens, H., Bochud, M., Braillard, L., Constandache, M., Jutzet, J.-M., 2010. Le gisement à faune Würmienne du Bärenloch (Préalpes fribourgeoises). Bulletin de la Société fribourgeoise des sciences naturelles SFSN 99: 1-22.

* Brinkman, D.B., Densmore, M.J., Joyce, W.G. 2010. “Macrobaenidae” (Testudines: Eucryptodira) from the Late Paleocene (Clarkforkian) of Montana and the taxonomic treatment of “Clemmys” backmani. Bulletin of the Peabody Museum of Natural History 51:147-155.

Bocherens, H. 2010. Isotopic Analysis of Keratin—Information about Living Environment and Nutrition. in: (A. Wieczorek, M. Tellenbach & W. Rosendahl, Eds) Mumies – The dream of eternal life, pp. 232-235.

* Campos, P.F., Kristensen, T., Orlando, L., Sher, A., Kholodova, M.V., Götherström, A., Hofreiter, M., Drucker, D.G., Kosintsev, P., Tikhonov, A., Baryshnikov, G.F., Willerslev, E., Gilbert, M.T.P. 2010. Ancient DNA sequences point to a large loss of mitochondrial genetic diversity in the saiga antelope (Saiga tatarica) since the Pleistocene. Molecular Ecology 19, 4863–4875. pdf

Drucker, D., Bocherens, H. 2010. Analyses isotopiques du carbone (13C) du collagene des restes de faune de Reute-Schorrenried. In: Fundberichte aus Baden-Württemberg. 31, pp. 89-95.

* Drucker, D.G., Hobson, K.A., Ouellet, J.-P., Courtois, R. 2010. Influence of forage preferences and habitat use on 13C and 15N abundance in wild caribou (Rangifer tarandus caribou) and moose (Alces alces) from Canada. Isotopes in Environmental and Health Studies 46,107-121. pdf

* Joyce, W.G. 2010. Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (Reptilia, Testudines). Bulletin of Zoological Nomenclature 67:78.

* Joyce, W.G., Lyson, T.R. 2010. A neglected lineage of North American turtles fills a major gap in the fossil record. Palaeontology 53:241-248.

* Joyce, W.G., Lyson, T.R. 2010. Pangshura tatrotia, a new species of pond turtle (Testudinoidea) from the Pliocene Siwaliks of Pakistan. Journal of Systematic Palaeontology 8:449-458.

* Lyson, T.R., Bever, G.S., Bhullar, B.-A.S., Joyce, W.G., Gauthier, J.A. 2010. Transitional fossils and the origin of turtles. Biology Letters 6:830-833.

* Lyson, T.R., Joyce, W.G., 2010. A new baenid turtle from the Late Cretaceous (Maastrichtian) Hell Creek Formation of North Dakota and a preliminary taxonomic revision of Cretaceous Baenidae. Journal of Vertebrate Paleontology 30:394-402.

* Pushkina, D., Bocherens, H., Chaimanee, Y., Jaeger, J-J. 2010. First dietary and paleoenvironmental reconstructions from the late Middle Pleistocene Snake cave in northeastern Thailand using stable carbon isotopes. Naturwissenschaften 97: 299-309. DOI 10.1007/s00114-009-0642-6

* Rabi, M., Joyce, W.G., Wings, O. 2010. A review of the Mesozoic turtles of the Junggar Basin (Xinjiang, Northwest China) and the paleobiogeography of Jurassic to Early Cretaceous Asian testudinates. Palaeobiodiversity and Palaeoenvironments 90:259-273.

* Stiller, M., Baryshnikov. G., Bocherens, H., Grandal d’Anglade, A., Hilpert, B., Münzel, S.C., Pinhasi, R., Rabeder, G., Rosendahl, W., Trinkaus, E., Hofreiter, M., Knapp, M. 2010. Withering away – 25,000 years of genetic decline preceded cave bear extinction. Molecular Biology and Evolution 27(5): 975-978. doi: 10.1093-/molbev/msq083 pdf media report 1 media report 2


*Audouze F., Drucker D.G., F. Valentin (2009) - Nouvelles données chronologiques, biologiques et alimentaires sur l'enfant mésolithique de Verberie - Le Buisson Campin (Oise), Bulletins et Mémoires de la Société d'Anthropologie de Paris 21 (1-2), p. 5-18. pdf
* Bocherens, H. 2009. Dental microwear of cave bears: the missing temperate/boreal vegetarian “carnivore”. PNAS 106(48), doi:10.1073/pnas.0910368106

* Bocherens, H. 2009. Neanderthal dietary habits: Review of the isotopic evidence. in: (J.-J. Hublin & M.P. Richards, Eds.) The Evolution of Hominid Diets: Integrating approaches to the study of Palaeolithic subsistence, Vertebrate Paleobiology & Paleoanthropology, Springer, Dordrecht, pp. 241-250.

* Bocherens, H. 2009. Possible freshwater resource consumption by the earliest directly dated European modern humans: Implications for direct radiometric dating. PNAS 106(41), doi:10.1073/pnas.0909392106

Bocherens, H. 2009. Sage mir, was du isst … Verräterische Isotope. In : (B. Theune-Großkopf & N. Conard, Eds) Eiszeit Kunst und Kulltur Archäologisches Landesmuseum Baden-Württemberg, Konstanz, Thorbecke p. 212.

* Bocherens, H., Jacques, L., Ogle, N., Moussa, I., Kalin, R., Vignaud, P. & Brunet, M. 2009. Reply to the comment by A. Zazzo, W.P Patterson and T.C. Prokopiuk on “Implications of diagenesis for the isotopic analysis of Upper Miocene large mammalian herbivore tooth enamel from Chad” By Jacques et al. (2008). Palaeogeography, Palaeoclimatology, Palaeoecology 266, 200-210.” Palaeogeography, Palaeoclimatology, Palaeoecology 277: 269-271 doi:10.1016/j.palaeo.2009.02.003

Drucker, D. G. & Bocherens, H. 2009. Carbon stable isotopes of mammal bones as tracers of canopy development and habitat use in temperate and boreal contexts. In: (J.D. Creighton & P.J. Roney, ed.) Forest Canopies: Forest Production, Ecosystem Health, and Climate Conditions Nova Science Publishers, Inc. ISBN 978-1-60741-457-5, 103-109.

* Drucker, D. G., Bocherens, H. & Billiou, D. 2009. Quelle valence écologique pour les rennes et autres cervidés dans les Alpes du Nord et le Jura ? Résultats de l'analyse des isotopes stables (13C, 15N) du collagène. In: (G. Pion Ed.) La fin du Paléolithique supérieur dans le nord, l'est de la France et les régions limitrophes, Mémoire de la Société préhistorique française 50, 73-86.

* Drucker, D. G., Bridault, A., Iacumin, P. & Bocherens, H. 2009. Bone stable isotopic signatures (15N, 18O) as tracer of temperature variation in Lateglacial and early Holocene: case study of red deer from Rochedane site in French Jura. Geological Journal 44: 593–604. DOI: 10.1002/gj.1170

* Joyce, W.G., Lucas, S.G., Scheyer, T., Heckert, A., Hunt., A.P. 2009. A thin-shelled reptile from the Late Triassic of North America and the origin of the turtle shell. Proceedings of the Royal Society of London, Series B, Biological Sciences 276:507-513.

* Joyce, W.G., Revan, A., Lyson, T.R., Danilov, I.G. 2009. Two new plastomenine softshell turtles from the Paleocene of Montana. Bulletin of the Peabody Museum of Natural History 50:307-325.

* Lyson, T.R., Joyce, W.G. 2009. A new species of Palatobaena (Testudines: Baenidae) and a maximum parsimony and Bayesian phylogenetic analysis of Baenidae. Journal of Paleontology 83:457-470.

* Lyson, T.R., Joyce, W.G. 2009. A revision of Plesiobaena (Testudines: Baenidae) and an assessment of baenid ecology across the K/T boundary. Journal of Paleontology 83:833-853.

* Manzano, A.S., Noriega, J.I., Joyce, W.G. 2009. The tropical tortoise Chelonoidis denticulata (Testudines: Testudinidae) from the late Pleistocene of Argentina and its paleoclimatological implications. Journal of Paleontology 83:974-979.

* Semal, P., Rougier, H., Crevecoeur, I., Jungels, C., Flas, D., Hauzeur, A., Maureille, B., Germonpré, M., Bocherens, H., Pirson, S., Cammaert, L., De Clerck, N., Hambuchen, A., Higham, T., Toussaint, M., van der Plicht, J. 2009. New data on the Late Neandertals: direct dating of the Belgian Spy fossils. American Journal of Physical Anthropology 138: 421-428. doi: 10.1002/ajpa.20954

* Tillier, A.-M., Mester, Z., Bocherens, H., Henry-Gambier, D., Pap, I. 2009. Direct dating of the “Gravettian” Balla child skeleton from Bükk Mountains (Hungary). Unexpected results. Journal of Human Evolution 56: 209-212. doi: 10.1016/j.jhevol.2008.09.005 pdf


* Bocherens, H., Drucker, D. G., Billiou, D., Geneste, J.-M. & Kervazo, B. 2008. Grotte Chauvet (Ardèche, France): a "natural experiment" for bone diagenesis in karstic context. Palaeogeography, Palaeoclimatology, Palaeoecology 266: 220-226. doi: 10.1016/j.palaeo.2008.03.023. pdf

* Bocherens, H. & Rousseau, L. 2008. Chapitre 8 : Le cadre environnemental des occupations humaines. Isotopes sur grands mammifères et plancher stalagmitique. in: (M.-H. Moncel, Ed.) Le site de Payre, Mémoire de la Société préhistorique française, 46, pp. 113-122.

* Coulson, A., Kohn, M., Shirley, M. Joyce, W.G. Barrick, R. 2008. Phosphate-oxygen isotopes from marine turtle bones: Ecologic and paleoclimatic applications. Palaeogeography, Palaeoclimatology, Palaeoecology 264:78-84.

* Döppes, D., Rosendahl, W., Pacher, M., Imhof, W., Dalmeri, G. & Bocherens, H. 2008. Stabile Isotopenuntersuchungen an spätglazialen und holozänen Braunbärenfunden aus Höhlen im Alpenraum. In: (T. Bürgin, Ed.) The 14th International Cave Bear Symposium at Appenzell, Switzerland, 18th-22nd of September 2008 Stalactite 58(2): 64-66. pdf

* Drucker, D. G., Bridault, A., Hobson, K. A., Szuma, E. & Bocherens, H. 2008. Can carbon-13 abundances in large herbivores track canopy effect in temperate and boreal ecosystems? Evidence from modern and ancient ungulates. Palaeogeography, Palaeoclimatology, Palaeoecology 266: 69-82. doi: 10.1016/j.palaeo.2008.03.020 pdf

Drucker D.G., Henry-Gambier D., Lenoir M. 2008. Nouvelles données chronologiques sur l'enfant de la grotte de Fauroux (Lugasson, Gironde) [New chronological data on the child of Fauroux I cave (Lugasson, Gironde)]. Paléo 20, 55-58. pdf

* Faux, C., Joyce, W.G. 2008. Connecticut's Pleistocene horse reassigned to the nineteenth century. Bulletin of the Peabody Museum of Natural History 49:131-132.

* Jacques, L., Ogle, N., Moussa, I., Kalin, R., Vignaud, P., Brunet, M. & Bocherens, H. 2008. Implications of diagenesis for the isotopic analysis of Upper Miocene large mammalian herbivore tooth enamel from Chad. Palaeogeography, Palaeoclimatology, Palaeoecology 266: 200-210. doi: 10.1016/j.palaeo.2008.03.040 pdf

* Joyce, W.G., Faux, C. 2008. “Chelys patagonica” from the Miocene of Argentina: Not lost, not valid, not a chelid turtle. Ameghiniana 12:801-803.

* Münzel, S.C., Hofreiter, M., Stiller, M., Conard, N.J. & Bocherens, H. 2008. Neue Ergebnisse zur Paläobiologie der Höhlenbären auf der Schwäbischen Alb (Chronologie, Isotopie und Paläogenetik). Stalactite 58(2): 27-30.

* Schulting, R., Blockey, S., Bocherens, H., Drucker, D. G. & Richards, M. 2008. Stable carbon and nitrogen isotope analysis on human remains from the Early Mesolithic site of La Vergne (Charente-Maritime, France). Journal of Archaeological Science 35: 763-772. doi:10.1016/j.jas.2007.06.008 pdf

* Valentin, F., Sand, C., Le Goff, I. & Bocherens, H. 2008. An early first millenium AD burial from the Naselala site, Cikobia-I-Ra Island (North-East Fiji). In: (D.J. Addison & C. Sand, Eds) Recent advances in the Archaeology of the Fidji/West-Polynesia region, University of Otago Studies in Prehistoric Anthropology, No. 21, Dunedin, pp. 45-56.

* Zelenitsky, D.K., Therrien, F., Joyce, W.G., Brinkman, D.B. 2008. First fossil gravid turtle provides insight into the evolution of reproductive traits in turtles. Biology Letters 4:715-718.