Institut für Evolution und Ökologie

List of publications

Evolutionary Biology of Invertebrates


Spiessberger, E. L., Newton, A. F., Thayer, M. K., Betz, O. (2024): Comparative morphology of the feeding apparatus of Staphylinine beetles (Coleoptera: Staphylinidae). Arthropod Systematics & Phylogeny 82: 267-303.

Wegst, U.G.K., Cloetens, P., Betz, O. (2024): Desert locusts (Schistocerca gregaria) feed with self-sharpening, scissor-like mandibles. Interface Focus 14 (2). 20230069.


van Nieuwenhoven, RW., Drack, M., Gebeshuber, IC. (2023): Engineered Materials: Bioinspired “Good Enough” versus Maximized Performance. Advanced Functional Materials 2023:2307127 (1-9).

Haas-Renninger, M., Weber, J., Felske, I., Kimmich, T., Csader, M., Betz, O., Krogmann, L., & Steidle, J. L. M. (2023): Microhymenoptera in roadside verges and the potential of arthropod-friendly mowing for their preservation. Journal of Applied Entomology, 00, 1–10.

Eggs, B.Fischer, S.Csader, M., Mikó, I., Rack, A., Betz, O. (2023): Terebra steering in chalcidoid wasps. Frontiers in Zoology 20, 26.

Zippel, A., Cao, Q., Betz, O. (2023): Morphology of the abdominal segmental glands and spinning behaviour of Stenus larvae (Coleoptera, Staphylinidae), Arthropod Structure and Development 75, 101286

Greb, I., Eggs, B. (2023): Die "Nosferatu-Spinne“ (Zoropsis spinimana): Wie gefährlich ist sie wirklich? Rettungsdienst, 46 (6), 572–575.

von Berg, L., Frank, J., Sann, M., Betz, O., Steidle, J.L.M., Böttinger, S. (2023): Insect-  and spider-friendly mowing technology in grassland – overview and evaluation, Landtechnik, 78 (2), 80-96.

Bruchhof, L., Axt, M., Betz, O. (2023): Untersuchungen zur Insektenvielfalt am Tübinger Bergfriedhof, Mitt. Ent. Ver. Stuttgart, 57 (1/2), 91-102

Yavorskaya, M. I., Jałoszyński, P., & Beutel, R. G. (2023): A unique case of commensalism: The beaver beetle Platypsyllus castoris (Leiodidae, Coleoptera) and its morphological adaptations. Journal of Morphology, 284(1), e21532.


Schädel, M., Yavorskaya, M., & Beutel, R. (2022): The earliest beetle† Coleopsis archaica (Insecta: Coleoptera)–morphological re-eva luation using Reflectance Transformation Imaging (RTI) and phylogenetic assessment. Arthropod Systematics & Phylogeny, 80, 495-510.

Drack, M., Betz, O. (2022): A technomorphic conceptualisation of biological ‘constructions’ and their evolution, Vertebrate Zoology, 72,  839–855,

Olmi, M., Eggs, B., Capradossi, L., van de Kamp, T., Perkovsky, E. E., Guglielmino, A., Vasilenko, D. V. (2022): A new species of Bocchus from upper Eocene Rovno amber (Hymenoptera, Dryinidae). Journal of Hymenoptera Research 92, 257–272.

Betz, O., Kimmich, T., Csader, M., Spinner, F., Steidle, J. (2022): Einsatz von Mähmaschinen mit schonender Wirkung auf die biologische Vielfalt zur Pflege von Straßenbegleitflächen am Beispiel des Grünpflegekopfs ECO 1200 plus® von MULAG, Natur und Landschaft, 97 (9/10), 455-461.

Stocker, B., Barthold, S., Betz, O. (2022): Mouthpart ecomorphology and predatory behaviour in selected rove beetles of the "staphylinine group" (Coleoptera: Staphylinidae: Staphylininae, Paederinae), insects, 13(8), 667;

Wanieck, K., Hamann, L., Bartz, M., Uttich, E., Hollermann, M., Drack, M., Beismann, H. (2022): Biomimetics linked to classical product development: An interdisciplinary endeavor to develop a technical standard. Biomimetics 7, 36.

Jedicke, E., Aufderheide, U., Bergmeier, E., Betz, O., Brunzel, S, Eckerter, P., Kirmer, A., Klatt, M., Kraft, M., Lukas, A., Mann, S., Mody, K., Schenkenberger, J., Schwenninger, H., Settele, J., Steidle, J.L.M., Tischew, S., Welk, E., Wolters, V., Worm, R. (2022): Gebietseigenes Saatgut – Chance oder Risiko für den Biodiversitätsschutz? Ein Thesenpapier zur Umsetzung des § 40 BNatSchG, Naturschutz und Landschaftsplanung, 54 (04), 12-21,

Steidle, J.L.M., Kimmich, T., Csader, M., Betz, O. (2022): Negative impact of roadside mowing on arthropod fauna and its reduction with ‘arthropod-friendly’ mowing technique, Journal of Applied Entomology,

van de Kamp, T., Mikó, I., Staniczek, A. H., Eggs, B., Bajerlein, D., Faragó, T., Hagelstein, L., Hamann, E., Spiecker, R., Baumbach, T., Janšta, P., Krogmann, L. (2022): Evolution of flexible biting in hyperdiverse parasitoid wasps. Proceedings of the Royal Society B: Biological Sciences 289(1967), 20212086.


Gaenshirt, P.Beyrich, M., Betz, O. (2021): Biological aspects of the larva of Sepedophilus testaceus (Coleoptera: Staphylinidae, Tachyporinae): morphology of the posterior abdomen and prey-capture behaviour. Journal of Natural History 55(31-32), 1999-2016.

Gebhardt, H., Beaver, R. A., Allgaier, Ch. (2021): Three new species of Scolytoplatypus Schaufuss from China, and notes on the movement and functions of the prosternal processes (Coleoptera: Curculionidae: Scolytinae).

Günther, A., Drack, M., Monod, L., Wirkner, C.S. (2021): A unique yet technically simple type of joint allows for the high mobility of scorpion tails. Journal of the Royal Society Interface 18: 20210388.
Featured by the New York Times, accessed 9 September 2021

Csader, M., Mayer, K., Betz, O., Fischer, S.Eggs, B. (2021): Ovipositor of the braconid wasp Habrobracon hebetor: structural and functional aspects, Journal of Hymenoptera Research, 83, 73–99.

Tucker, C., Yang, X., Lehrecke, A., Skoury, L., Aust, M., Holtmann, F., Körner, A., Betz, O. (2021): Bending active plates with tensile elements: a biomimetic approach inspired by the Australian Curly Leaf Spider (Phonognatha graeffae). Proceedings of the IASS Annual Symposium 2020/21 and the 7th International Conference on Spatial Structures Inspiring the Next Generation, 1-12.


Drack, M., Berger, A., Ettinger, B., Gebeshuber, I.C. (2020): 3D-printed facet optics: Novel adjustable technical optics inspired by compound eyes. Frontiers in Materials 7:199.

Drack, M., Betz, O., Nebelsick, J.H. (2020): Konstruktionslehre, Bionik und phylogenetische Aspekte, in: Werneburg, I. & Betz, O. (Hrsg.) (2020): Phylogenie, Funktionsmorphologie und Bionik, Scidinge Hall, 27-38.

Barthold, S. & Betz, O. (2020): The predatory behaviour of some Central European rove beetles (Coleoptera: Staphylinidae: Staphylininae, Paederinae), Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 22, 173-178.

Tull T., Henn F., Betz, O.Eggs B. (2020). Structure and function of the stylets of hematophagous Triatominae (Hemiptera: Reduviidae), with special reference to Dipetalogaster maxima. Arthropod Structure & Development 52, 100952.

Mohr T., Fischer S. (2020) Ultrastructural evidence for the origin of the sub-retinal pigment shield in the compound eye of Drosophila melanogaster. J. Morph. 281: 802-807.

Betz O., Srisuka, W., Puthz, V.  (2020). Elevational gradients of species richness, community structure, and niche occupation of tropical rove beetles (Coleoptera: Staphylinidae: Steninae) across mountain slopes in Northern Thailand. Evolutionary Ecology 34, 193–216.

Werneburg I., Betz O. (2020) Phylogenie, Funktionsmorphologie und Bionik. Schriften zum 60. Phylogenetischen Symposium in Tübingen. Scidinge Hall Verlag Tübingen, 332 pages.


Betz O., Eggs B., Henn F., Birkhold A., Röhrle O. (2019): Movement without joints: (how) does it work? In: Knippers J., Schmid U., Speck T. (eds.) (2019) Biomimetics for Architecture. Learning from Nature, Birkhäuser, 22-31.

Betz O., Heethoff M., Garamszegi L.Z., Koerner L. (2019): The beetles with the protrusible tongue: Integration of form, function, and ecology in the predatory rove beetles Stenus spp., Integrative and Comparative Biology 59, Issue Supplement_1, e17,

De Bruyn G., Betz O., Drack M., Limpinsel M., Nebelsick J. (2019): Functionalist, organic, and biomimetic architecture. In: Knippers J., Schmid U., Speck T. (eds.) (2019) Biomimetics for Architecture. Learning from Nature, Birkhäuser, 172-179.

Drack M. (2019) Towards a formalisation of biomimetics via engineering design concepts. Pages 92–102 in: Bionik: Patente aus der Natur, Tagungsbeiträge, 9. Bionik-Kongress, edited by Kesel AB, Zehren D. Bremen: Bionik-Innovations-Centrum (B-I-C). From the talk at the conference “Bionik-Kongress 2018 ‘Patente aus der Natur’”. 26.–27.10.2018, Bremen (Germany). Organizer: Antonia B. Kesel, Gesellschaft für Technische Biologie und Bionik (GTBB), Hochschule Bremen, Bionik-Innovations-Centrum.

Fischer S., Lu Z., Meinertzhagen I.A. (2019) Three-dimensional ultrastructural organization of the ommatidium of the minute parasitoid wasp Trichogramma evanescens. Arthropod Structure and Development 48:35-48.

Mohr T., Meinertzhagen I.A., Fischer S. (2019) Novel type of sub‐retinal pigment shield in the miniaturized compound eye of Trichogramma evanescens. J Comp Neurol 

Schnee, L., Sampalla, B., Müller, J., Betz, O. (2019), A comparison of tarsal morphology and traction force in the two burying beetles Nicrophorus nepalensis and Nicrophorus vespilloides (Coleoptera, Silphidae), Beilstein Journal of Nanotechnology, 10, 47-61. doi:10.3762/bjnano.10.5

Zobec, M., Betz, O., Unterweger, P.A, (2019): Perception of urban green areas associated with sociodemographic affiliation, structural elements, and acceptance stripes, urban science 4, 9, 1-40, doi:10.3390/urbansci4010009


Bergmann, P.A., Glöckner, N.,  Richter, S., Betz, O. (2018): Morphology of hindwing veins in the shield bug Graphosoma italicum (Heteroptera: Pentatomidae), Arthropod Struct. Dev., 47 (4), 375-390.

Bergmann, P., Laumann, M., Norton, R.A., Heethoff, M. (2018). Cytological evidence for automictic thelytoky in parthenogenetic oribatid mites (Acari, Oribatida): Synaptonemal complexes confirm meiosis in Archegozetes longisetosus. Acarologia 58: 342-356.

Betz, O., Albert, K., Boley, M., Frenzel, M., Gerhardt, H., Grunwald, I., Hartwig,A., Kleemeier, M., Maurer, A., Neuenfeldt, M., Rischka, K., Sampalla, B., Schmitt, C., Speidel, M., Steiner, M., Verheyden, N., Vogt, M. (2018): Struktur und Funktion des tarsalen Haftsystems der Madagaskar-Fauchschabe Grompadorhina portentosa (Blattodea), Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 21, 159-164.

Betz, O., Irmler, U. & Klimaszewski, J. (eds.) (2018): Biology of rove beetles (Staphylinidae) - Life history, evolution, ecology and distribution, Springer, 351 pp.

Betz, O., Koerner, L., Dettner, K. (2018); The biology of Steninae, in: Betz, O., Irmler, U. & Klimaszewski, J. (eds.), Biology of rove beetles (Staphylinidae) - Life history, evolution, ecology and distribution, Springer, 229-283.

Drack M., Limpinsel M., de Bruyn G., Nebelsick J.H., Betz O. (2018) Towards a theoretical clarification of biomimetics using conceptual tools from engineering design. Bioinspiration & Biomimetics13: 016007.

Eggs B., Birkhold A.I., Röhrle O., Betz O. (2018): Structure and function of the musculoskeletal ovipositor system of an ichneumonid wasp. BMC Zoology 3, 12.

Eggs B., Birkhold A. I., Röhrle O., Betz O. (2018). The musculoskeletal ovipositor system of an ichneumonid wasp: structural and functional aspects. Mitteilungen des Entomologischen Vereins Stuttgart 53 (1), 34–36. 

Irmler, U., Klimaszewski, J. & Betz, O. (2018) Introduction to the biology of rove beetles, in: Betz, O., Irmler, U. & Klimaszewski, J. (eds.), Biology of rove beetles (Staphylinidae) - Life history, evolution, ecology and distribution, Springer, 1-4.

Krogmann, L., Betz, O., Geldmann, J., Goulson, D., Menzel, R., Riecken, U., Ruther, J., Schwenninger, H. R., Sorg, M., Steidle, J., Tscharntke, T., Wägele, W. (2018) Neun-Punkte-Plan gegen das Insektensterben – Die Perspektive der Wissenschaft. Entomologische Zeitschrift • Schwanfeld • 128 (4), 247-249.

Nadein, K. & Betz, O. (2018): Jumping mechanisms and performance in beetles. II. Weevils (Coleoptera: Curculionidae: Rhamphini), Arthropod Struct. Dev., 47, 131-143.

Qarony W., Hossain M.I., Dewan R., Fischer S., Meyer- Rochow V.B., Salleo A., Knipp D., Tsang Y.H. (2018) Approaching perfect light incoupling in perovskite and silicon thin film solar cells by moth eye surface textures. Advanced Thory and Simulations 1 (8) 1800030.

Sampalla, B., Eggs, B., Betz, O. (2018): Bending the sting: joint-free movement principles in the ovipositor of the parasitoid wasp Leptopilina heterotoma (Figitidae), Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 21, 171-174.

Schierbaum,N., Hack M., Betz, O., Schäffer, T.E. (2018): Macro-SICM: A Scanning Ion Conductance Microscope for Large-Range Imaging, Analytical Chemistry, 90 (8), 5048-5054.

Tinello D, Winkler H, Drack M, Gebeshuber IC, Bevanda-Rauch D. 2018. Bio-inspirierte Fabriklayouts – Natur-Modelle für einen effizienten Materialfluss. Hebezeuge Fördermittel 03/2018:34–36. (ISSN 0017-9442)

Unterweger, P.A., Klammer, J., Unger, M., Betz, O. (2018): A winter-adapted green space management for insect conservation: a hitherto neglected tool for biodiversity protection, BioRisk 13, 1-29.

Werneburg, I., Betz, O. (2018): Das 60. Phylogenetische Symposium in Tübingen: Funktionsmorphologie und Bionik - Einführung. In: Werneburg I, Betz O (Hrsg.). Programm und Abstracts. 23. - 25. November 2018. Scidinge Hall Verlag Tübingen, 38 Seiten, ISBN: 978-3-947020-07-2 (digital auf: TOBIAS-lib, doi:

Werneburg, I., Betz O. (2018, Hrsg.): 60. Phylogenetische Symposium. Funktionsmorphologie und Bionik. Programm und Abstracts. 23. - 25. November 2018. Scidinge Hall Verlag Tübingen, S. 3-15.


Betz, O., Eggs, B., Henn, F., Birkhold, A., Röhrle, O. (2017) Bewegung ohne Gelenke: (Wie) geht das? In: Knippers, J., Schmid, U., Speck, T. Baubionik – Biologie beflügelt Architektur (Stuttgarter Beiträge zur Naturkunde, Serie C, Band 82), Staatliches Museum für Naturkunde Stuttgart und Gesellschaft zur Förderung des Naturkundemuseums in Stuttgart e.V., 20-29.

Betz, O., Frenzel, M., Steiner, M., Vogt, M., Kleemeier, M., Hartwig, A., Sampalla, B., Rupp, F., Boley, M., Schmitt, C. (2017): Adhesion and friction of the smooth attachment system of the cockroach Gromphadorhina portentosa and the influence of the application of fluid adhesives, Biology Open, 6, 589-601, DOI:10.1242/bio.024620

Born, L., Körner, A., Schieber, G., Westermeier, A.S., Poppinga, S., Sachse, R., Bergmann, P., Betz, O., Bischoff, M., Speck, T., Knippers, J., Milwich, M., Gresser, G.T. (2017): Fiber-reinforced plastics with locally adapted stiffness for bio-inspired hingeless, deployable architectural systems. In: Herrmann A (ed.) 21st Symposium on Composites, Bd. 742: Trans Tech Publications (Key Engineering Materials), p. 689–696.

de Bruyn, G., Betz, O., Drack, M., Limpinsel, M., Nebelsick, J.N. (2017) Funktionalistische, organische und bionische Architektur, In: Knippers, J., Schmid, U., Speck, T. Baubionik – Biologie beflügelt Architektur (Stuttgarter Beiträge zur Naturkunde, Serie C, Band 82), Staatliches Museum für Naturkunde Stuttgart und Gesellschaft zur Förderung des Naturkundemuseums in Stuttgart e.V., 130-137.

Drack M. (2017) An Affinity to Theories in Biology. Pages 79–86 in: Philosophy of Systems Biology: Perspectives from Scientists and Philosophers, edited by Green S. Cham (Switzerland): Springer. (DOI: 10.1007/978-3-319-47000-9_7) (ISBN: 978-3-319-46999-7)

Drack, M., Betz, O. (2017). The basis of theory building in biology lies in the organism concept: a historical perspective on the shoulders of three giants. Organisms. Journal of Biological Sciences 1: 69–82. DOI: 10.13133/2532-5876_2.11

Drack, M., de Bruyn, G., Limpinsel, M., Nebelsick, J., Betz, O. (2017): Towards a theoretical clarification of biomimetics using conceptual tools from engineering design, Bioinspiration and Biomimetics, 13 (1), 016007

Fischer S., Lu Z., Meinertzhagen I.A. (2017) From two to three dimensions: The importance of the third dimension for evaluating the limits to neuronal miniaturization in insects. J Comp Neurol. (526 (4):653-662).

Koerner, L., Garamszegi, L.Z., Heethoff, M., Betz, O. (2017): Divergent morphologies of adhesive predatory mouthparts of Stenus species (Coleoptera: Staphylinidae) explain differences in adhesive performance and resulting prey-capture success, Zool. J. Linn. Soc. 181: 500-518.

Schieber, G., Born, L., Bergmann, P., Körner, A., Mader, A., Saffarian, S., Betz, O., Milwich, M., Gresser, G.T., Knippers, J. (2017): Hindwings of insects as concept generator for hingeless foldable shading systems, Bioinspiration and Biomimetics, 13, 016012.

Schmelzle, S., Heethoff, M., Heuveline, V., Lösel, P., Becker, J., Beckmann, F., Schluenzen, F., Hammel,J.U., Kopmann, A., Mexner, W., Vogelgesang,M., Jerome, N.T., Betz, O., Beutel, R., Wipfler,B., Blanke, A., Harzsch, S., Hörnig, M., Baumbach, T., van de Kamp, T. (2017): The NOVA project: maximizing beam time efficiency through synergistic analyses of SRμCT data, Proc. SPIE 10391, Developments in X-Ray Tomography XI, 103910P (19 September 2017); doi: 10.1117/12.2275959;

Schmitt, C. & Betz, O. (2017): Morphology and ultrastructure of the tarsal adhesion organs of the Madagascar hissing cockroach Gromphadorhina portentosa, Cell and Tissue Research, 370, 243-265.

Speidel, M. W., Kleemeier, M., Hartwig, A., Rischka, K., Ellermann, A., Daniels, R. & Betz, O. (2017): Structural and tribometric characterization of biomimetically inspired synthetic "insect adhesives", Beilstein Journal of Nanotechnology, 8, 45-63

Tinello D, Boley M, Drack M, Gebeshuber IC, Winkler H. 2017. Bionik in der Logistik – umsetzbares Potential! Pages 125–139 in: Logistisches Produktionsmanagement: 4. Wissenschaftlicher Industrielogistik-Dialog in Leoben (WiLD), edited by Zsifkovits HE, Altendorfer-Kaiser S. Berlin: epubli. ISBN 9783745016314

Unterweger, P.A., Rieger, C., Betz, O. (2017): The influence of urban lawn mowing regimes on heteropteran diversity, Heteropteron, 48, 7-21.

Unterweger, P., Schrode, N., Potthast, T., Betz, O. (2017): Eine Problemfeldanalyse des urbanen Naturschutzes. Korrespondenz und Medienresonanz zur Arbeit der Initiative „Bunte Wiese – für mehr Artenvielfalt auf öffentlichem Grün“ in Tübingen, Naturschutz und Landschaftsplanung, 49 (8), 245-251.


Betz O., Birkhold A., Caliaro M., Eggs B., Mader A., Knippers J., Röhrle O., Speck O. (2016). Adaptive stiffness and joint-free kinematics: actively actuated rod-shaped structures in plants and animals and their biomimetic potential in architecture and engineering. In: Knippers J., Nickel K. G. and Speck T. (eds.), Biomimetic Research for Architecture and Building Construction: Biological Design and Integrative Structures (pp. 135–167), Biologically-Inspired Systems, vol. 8. Springer International Publishing.

Betz, O., Verheyden, A.N., Maurer, A., Schmitt, C., Braun, J., Kowalik, T., Grunwald, I., Hartwig, A. & Neuenfeldt, M. (2016): First protein and peptide characterization of the tarsal adhesive secretions in the desert locust, Schistocerca gregaria, and the Madagascar hissing cockroach, Gromphadorhina portentosa, Insect Molecular Biology, 25 (5), 541-549. doi: 10.1111/imb.12241

Bruyn, G. de, Betz, O., Nebelsick, J.H., Drack, M., Limpinsel, M. (2016): Making life "visible": Organism concepts in biology and architecture as the basis for an interdisciplinary synopsis of constructional biomimetics, in: Knippers, J., Nickel, K. & Speck, T. (Eds.): Biomimetic Research for Architecture and Building Construction. Biological Design and Integrative Structures. Springer, 379-394.

Drack M. (2016). On connections between properties in biology – How we learn about order in organisms. Systema4:48–53. (

Drack M. (2016) The layer model of Rupert Riedl. Pages 312–328 in: Aktuelle Bloch-Studien 2013–2014Münchener Schriften zur Design Science, Vol. 8, edited by Zimmermann RE. Aachen: Shaker. From the talk at the conference “Metaphysik der Emergenz – Philosophische Grundlegung im Spannungsfeld von Wissenschaft und Kunst”. 6.–8.6.2013, Ludwigshafen (Germany). Organizer: Rainer Zimmermann. (ISBN 978-3-8440-4782-0)

Gebeshuber I C, Drack M. (2016) Biomimetics: Biomimetics in nanotechnology. Pages 37–45 in: CRC Concise Encyclopedia of Nanotechnology, edited by Kharisov B I, Kharissova O V, Ortiz-Mendez U. Boca Raton (FL): CRC Press.

Gerhardt, H., Betz, O., Albert, K. & Lämmerhofer, M. (2016): Similarities, dissimilarities and classification of molecular (hydrocarbon) profiles of insect secretions in dependence of species, sex, and sampled body region, Journal of Chemical Ecology, 42, 725-738. DOI 10.1007/s10886-016-0718-7

Hartmann, K., Laumann, M., Bergmann, P., Heethoff, M., Schmelzle, S. (2016). Development of the synganglion and morphology of the adult nervous system in the mite Archegozetes longisetosus Aoki (Chelicerata, Actinotrichida, Oribatida). Journal of Morphology 277: 537-548.

Koerner, L. Braun, V. & Betz, O. (2016): The labial adhesive pads of rove beetles of the genus Stenus (Coleoptera: Staphylinidae) as carriers of bacteria, Entomologia Generalis, 36 (1), 33-41. doi: 10.1127/entomologia/2016/0308

Nadein, K. & Betz, O. (2016): Jumping mechanisms and performance in beetles. I: Flea beetles (Coleoptera: Chrysomelidae: Alticini), Journal of Experimental Biology, 219, 2015-2027. doi:10.1242/jeb.140533

Rack, A., Olbinado, M., Scheel, M., Schmitt, C., Betz, O., Garcia-Moreno, F., Banhart, J., Lhuissier, P., Just, X., Lhuissier, P., Gelet, J.-L-. Morse, J. (2016): Real-time X-ray microimaging using hard synchrotron radiation, The 31st International Congress on High-speed Imaging and Photonics, November 7-10, 2016, Osaka, JAPAN.

Wastian, L., Unterweger, P. A. & Betz, O. (2016): Influence of the reduction of urban lawn mowing on wild bee diversity (Hymenoptera, Apoidea), Journal of Hymenoptera Research, 49, 51-63.


Drack M. (2015) Ludwig von Bertalanffy’s organismic view on the theory of evolution. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution324B: 77–90. DOI: 10.1002/jez.b.22611 (invited)

Drack M., Pouvreau D. (2015) On the history of Ludwig von Bertalanffy’s “General Systemology”, and on its relationship to cybernetics – Part III: Convergences and divergences. International Journal of General Systems44: 523–571. DOI: 10.1080/03081079.2014.1000642

Eggs B., Wolff J. O., Kuhn-Nentwig L., Gorb S. N., Nentwig W. (2015). Hunting without a web: how lycosoid spiders subdue their prey. Ethology 121(12), 1166–1177.

Foelix R. F., Erb B., Eggs B. (2015). Morphologische Besonderheiten der Kräuseljagdspinne Zoropsis spinimana. ARACHNE 20(3), 4–14.

Gerhardt, H., Schmitt, C., Betz, O., Albert, K., Lämmerhofer, M. (2015): Contact solid-phase microextraction with uncoated glass and polydimethylsiloxane-coated fibers versus solvent sampling for the determination of hydrocarbons in adhesion secretions of Madagascar hissing cockroaches Gromphadorrhina portentosa (Blattodea) by gas chromatography-mass spectrometry, Journal of Chromatography A, 1388, 24-35.

Lang C., Koerner L., Betz O., Puthz V. & Dettner K. (2015): Phylogenetic relationships and chemical evolution of the genera Stenus and Dianous (Coleoptera: Staphylinidae). Chemoecology, 25 (1), 11-24.

Reitz M., Gerhardt H., Schmitt C., Betz O., Albert K. & Lämmerhofer M. (2015): Analysis of chemical profiles of insect adhesion secretions by gas chromatography-mass spectrometry, Analytica Chimica Acta, 854, 47-60.

Unterweger P., Braun A., Boess R., Koltzenburg M. & Betz O. (2015): Langfristige Etablierung extensiver Grünflächenpflege in Stadtgebieten. Vorstellung der Initiative „Bunte Wiese“ der Stadt Tübingen, DGaaE-Nachrichten 29 (2), 73-80.


Betz, O. (2014): Entstehung und evolutive Entfaltung der Insekten, in: Maier, W. & Werneburg, I. (Hrsg.), Schlüsselereignisse der organismischen Makroevolution. Scidinge Hall, 163-214.

Hiller, D. & Betz, O. (2014): Auswirkungen verschiedener Mahdkonzepte auf die Heuschreckenfauna städtischer Grünflächen. Untersuchungen auf Grünflächen in Tübingen. Naturschutz und Landschaftsplanung, 46 (8), 241-246.

Kricke, C. Bamann, T. & Betz, O. (2014): Einfluss städtischer Mahdkonzepte auf die Artenvielfalt der Tagfalter. Naturschutz und Landschaftsplanung, 46 (2), 052-058.

Schmitt, C., Rack, A. & Betz, O. (2014): Analyses of the mouthpart kinematics in Periplaneta americana (Blattodea, Blattidae) using Synchrotron X-ray cineradiography, Journal of Experimental Biology, 217, 3095-3107.

Unterweger, P., Braun, A., Koltzenburg, M. & Betz, O. (2014): Die Initiative Bunte Wiese – Für mehr Artenvielfalt auf öffentlichem Grün. In: Biodiversität und Klima – Vernetzung der Akteure in Deutschland X. Ergebnisse und Dokumentation des 10. Workshops. BFN-Skripten 357, 53-54.

Weide, D. & Betz, O. (2014): Kopfmorphologische Untersuchungen von Sporenfressern und Nicht-Sporenfressern bei Kurzflügelkäfern (Staphylinidae, Coleoptera) mittels Synchrotron-Mikrotomographie, Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 19, 173-178.

Weide, D., Thayer, M.K., Betz, O. (2014): Comparative morphology of the tentorium and hypopharyngeal-premental sclerites in sporophagous and non-sporophagous adult Aleocharinae (Coleoptera: Staphylinidae), Acta Zoologica, 95, 84-110.


Eggs B., Sanders D. (2013). Herbivory in spiders: the importance of pollen for orb-weavers. PLoS ONE 8(11), e82637.

Heethoff, M., Bergmann, P., Laumann, M., Norton, R. A. (2013). The 20th anniversary of a model mite: A review of current knowledge about Archegozetes longisetosus (Acari, Oribatida). Acarologia 53: 353-368.

Koerner, L., Laumann, M., Betz, O. & Heethoff, M. (2013): Loss of the sticky harpoon - COI sequences indicate paraphyly of Stenus with respect to Dianous (Staphylinidae, Steninae), Zoologischer Anzeiger, 252 (3), 337-347.

Unterweger, P., Ade, J., Braun, A., Koltzenburg, M., Kricke, C., Schnee, L. Wastian, L. & Betz, O. (2013): , Langfristige Etablierung extensiver Grünflächenpflege in Stadtgebieten. Die Initiative „Bunte Wiese“ der Stadt Tübingen. in Korn, H. & Feit, U. (Bearb.): Treffpunkt Biologische Vielfalt XII : interdisziplinärer Forschungsaustausch im Rahmen des Übereinkommens über die biologische Vielfalt, 89-94.

2012 und älter


Ade, J., Wolf-Schwenninger, K., Betz, O. (2012): Auswirkungen der Wiesenmahd auf verschiedene Käferarten ausgewählter städtischer Grünflächen Tübingens, Jahreshefte der Gesellschaft für Naturkunde in Württemberg, 168, 199-216.

Bergmann, P., Heethoff, M. (2012). Development of the internal reproductive organs in early nymphal stages of Archegozetes longisetosus Aoki (Acari, Oribatida, Tthypochthoniidae) as obtained by synchrotron X-ray microtomography (SR-µCT) and transmission electron microscopy (TEM). Soil Organisms 84: 459-470.

Bergmann, P., Heethoff, M. (2012). The oviduct is a brood chamber for facultative egg retention in the parthenogenetic oribatid mite Archegozetes longisetosus Aoki (Acari, Oribatida), Tissue & Cell 44: 342-350.

Heethoff, M. (2012). Regeneration of complex defensive oil-gland secretions and its importance for chemical defense in an oribatid mite. J. Chem. Ecol. 38: 1116-1123.

Heethoff, M., Raspotnig, G. (2012). Expanding the ‘enemy-free space’ for oribatid mites: Evidence for chemical defense of juvenile Archegozetes longisetosus against the rove beetle Stenus juno. Exp. Appl. Acarol. 56: 93-97.

Heethoff, M., Raspotnig, G. (2012). Investigating chemical communication in oribatid and astigmatid mites in bioassays - Pitfalls and suggestions. Soil Organisms 84: 409-421.

Heethoff, M., Raspotnig, G. (2012). Triggering chemical defense in an oribatid mite using artificial stimuli. Exp. Appl. Acarol. 56: 287-295.

Koerner, L., Gorb, S., Betz, O. (2012): Functional morphology and adhesive performance of the stick-capture apparatus of the rove beetles Stenus spp.(Coleoptera, Staphylinidae), Zoology, 115, 117-127.

Koerner, L., Gorb, S., Betz, O. (2012): Adhesive performance of the stick-capture apparatus of rove beetles of the genus Stenus (Coleoptera, Staphylinidae) toward various surfaces, Journal of Insect Physiology, 58, 155-163

Schmelzle, S., Norton, R. A., Heethoff, M. (2012). A morphological comparison of two closely related ptychoid oribatid mite species: Phthiracarus longulus and P. globosus (Acari: Oribatida: Phthiracaroidea). Soil Organisms 84: 431-443.

Van der Meijden, A., Langer, F., Boistel, R., Vagovic, P., Heethoff, M. (2012). Functional morphology and bite performance of raptorial chelicerae of camel spiders (Solifugae). J. Exp. Biol. 15: 3411-3418.


Alberti, G., Heethoff, M., Norton, R. A., Schmelzle, S., Seniczak, A., Seniczak, S. (2011). Fine structure of the gnathosoma of Archegozetes longisetosus Aoki (Acari: Oribatida, Trhypochthoniidae). J. Morphology 272: 1025-1079.

Betz, O. (2011): Artentstehung, natürliches Aussterben und anthropogene Krise der Biodiversität - Darwins Theorie aus heutiger evolutionsbiologischer Sicht, in: Engels, E.-M., Betz, O., Köhler, H.-R. & Potthast, T. (Hrsg.), Charles Darwin und seine Wirkung in Wissenschaft und Gesellschaft, Attempto, 89-119.

Engels, E.-M., Betz, O., Köhler, H.-R. & Potthast, T. (Hrsg.) (2011): Charles Darwin und seine Wirkung in Wissenschaft und Gesellschaft, Attempto, 291 pp.

Engels, E.-M., Betz, O., Köhler, H.-R. & Potthast, T. (2011): Charles Darwin und seine Bedeutung für die Wissenschaften – eine Einführung, in: Engels, E.-M., Betz, O., Köhler, H.-R. & Potthast, T. (Hrsg.): Charles Darwin und seine Wirkung in Wissenschaft und Gesellschaft, Attempto, 9-25.

Fernández, R., Bergmann, P., Almodóvar, A., Cosín, D., Heethoff, M. (2011). Ultrastructural and molecular insights into three populations of Aporrectodea trapezoides (Dugés, 1828) (Oligochaeta, Lumbricidae) with different reproductive modes. Pedobiologia 54: 281-290.

Heethoff, M., Koerner, L., Norton, R. A., Raspotnig, G. (2011). Tasty but protected - first evidence of chemical defense in oribatid mites, J. Chem. Ecol. 37: 1037-1043.

Heethoff, M., Laumann, M., Weigmann, G., Raspotnig, G. (2011). Integrative taxonomy: combining morphological, molecular and chemical data for species delineation in the parthenogenetic Trhypochthonius tectorum complex (Acari, Oribatida, Trhypochthoniidae), Front. Zool. 8:2.


Bergmann, P., Laumann, M., Heethoff, M. (2010). Ultrastructural aspects of vitellogenesis in Archegozetes longisetosus Aoki 1965 (Acari, Oribatida, Trhypochthoniidae). Soil Organisms 82: 193-208.

Betz, O. & Köhler, H.-R. (Hrsg.) (2008): Die Evolution des Lebendigen. Grundlagen und Aktualität der Evolutionslehre. Attempto, 304 pp.

Betz, O. (2010): Adhesive exocrine glands in insects: morphology, ultrastructure, and adhesive secretion, in: Byern, J. & Grunwald, I. (eds.): Biological adhesive systems. From nature to technical and medical application, Springer, 111-152.

Laumann, M., Bergmann, P., Norton, R. A., Heethoff, M. (2010).  First cleavages, preblastula and blastula in the parthenogenetic mite Archegozetes longisetosus (Acari, Oribatida) indicate holoblastic rather than superficial cleavage. Arthropod Struct. Dev. 39: 276.286.

Laumann, M., Norton, R. A., Heethoff, M. (2010). Acarine cleavage patterns and early embryology: Inconsistencies, artificial results and misinterpretation. Soil Organisms 82: 217-235.

Osterauer, R., Marschner, L., Betz, O., Gerberding, M., Sawasdee, B., Cloetens, P., Haus, N., Sures, B., Triebskorn, R., Köhler, H.-R. (2010): Turning snails into slugs: Induced body plan changes and formation of an internal shell. Evolution and Development, 12 (5), 474-483.

Rack, A., García-Moreno, F., Banhart, J., Schmitt, C., Betz, O., Cecilia, A., Ershov, A., Rack, T. & Zabler, S. (2010): On the possibilities of hard X-ray imaging with high spatio-temporal resolution using polychromatic synchrotron radiation, Journal of XRay Science and Technology 18, 429–441.

Schmelzle, S., Helfen, L., Norton, R. A., Heethoff, M. (2010). The ptychoid defensive mechanism in Phthiracarus longulus (Acari, Oribatida, Phthiracaroidea): Exoskeletal amd muscular elements. Soil Organisms 82: 253-273.

Weide, D., Thayer, M.K., Newton, A.F. & Betz, O. (2010): Comparative studies on the head of selected sporophagous and non-sporophagous Aleocharinae (Coleoptera: Staphylinidae): musculature and hypopharynx-prementum complex, Journal of Morphology 271, 910-931.

Wenk, P., Lucic, S. & Betz, O. (2010): Functional anatomy of the hypopharynx and the salivary pump in the feeding apparatus of the assassin bug Rhodnius prolixus (Stål 1858) (Heteroptera: Reduviidae), Zoomorphology 129, 225-234.


Bamann, T. & Betz, O. (2009). Die Libellen des NSG “Schaichtal” (Schönbuch) – Ergebnisse einer ökologisch orientierten Diplomarbeit, mercuriale – Libellen in Baden-Württemberg, 1-10.

Betz, O., Koerner, L., Gorb, S. (2009): An insect's tongue as the model for two-phase viscous adhesives? adhesion Adhesives & Sealants 3, 32-35.

Dieterich, A. & Betz, O. (2009): Elementsensitive Synchrotron-Mikrotomographie zur Darstellung von Zinkeinlagerungen in den Mandibeln ausgewählter Insekten, Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie, 17, 285-288.

Heethoff, M. (2009). Mit Röntgenstrahlen in den Mikrokosmos – Synchrotron-Röntgen-Mikrotomographie für nicht-invasive Visualisierung mikroskopischer Strukturen. BIOforum 32: 14-16.

Heethoff, M., Helfen, L., Norton, R. A. (2009) Description of Neoliodes dominicus n.sp. (Acari, Oribatida) from Dominican amber, aided by synchrotron X-ray microtomography. J. Paleont. 83: 153-159.

Heethoff, M., Norton, R. A. (2009). A new use of synchrotron X-ray microtomography (SR-µCT): three-dimensional biomechanical modeling of chelicerate mouthparts and calculation of theoretical bite forces. Inv. Biol. 128: 332-339.

Heethoff, M., Norton, R. A. (2009). Role of musculature during defecation in a particle-feeding arachnid, Archegozetes longisetosus (Acari, Oribatida). J. Morphology 270: 1-13.

Heethoff, M., Norton, R. A., Scheu, S., Maraun, M. (2009). Parthenogenesis in oribatid mites (Acari, Oribatida): evolution without sex. In: Schoen, I., Martens, K., van Dijk, P. (eds.) Lost Sex - The Evolutionary Biology of Parthenogenesis. Springer Press, pp 241-257.

Marschner, L., Osterauer, R., Betz, O.GerberdingM., Cloetens, P., Sures, B., Triebskorn, R., Köhler, H.-R. (2009): Platinum-induced internalization of shell formation in snails, Comparative Biochemistry and Physiology A, 154 (1), S18.

Rack, A., García-Moreno, F., Betz, O., Zabler, S., Schmitt, C., dos Santos Rolo, T., Ershov, A., Helfen, L., Banhart, J. & Baumbach, T. (2009): Synchrotron-based radioscopy with spatio-temporal micro-resolution using hard X-rays, Nuclear Science Symposium Conference Record, 2008. NSS '08. IEEE, 528-531, DOI: 10.1109/NSSMIC.2008.4775234.

Schmelzle, S., Helfen, L., Norton, R. A., Heethoff, M. (2009). The ptychoid defensive mechanism in Euphthiracaroidea (Acari: Oribatida): A comparison of muscular elements with functional considerations. Arthropod Struct. Dev. 38: 461-472.

Schmitt, C., Rack, A., Körner, L., Dieterich, A., Zabler, S., Betz, O. (2009): High-speed X-ray cineradiography for imaging the mouthpart kinematics of living insects, ANKA Highlights 2009, 25-26.

Weide, D. & Betz, O. (2009): Head morphology of selected Staphylinoidea (Coleoptera: Staphyliniformia) with an evaluation of possible groundplan features in Staphylinidae, Journal of Morphology 270, 1503-1523.

Wenk, P., Lucic, S. & Betz, O. (2009): Funktionelle Anatomie des Hypopharynx und der Speichelpumpe beim Saugapparat der Raubwanze Rhodnius prolixus (Stål 1858) (Heteroptera: Reduviidae), Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie, 17, 281-284.


Bergmann, P., Laumann, M., Cloetens, P., Heethoff, M. (2008). Morphology of the internal reproductive organs of Archegozetes longisetosus Aoki (Acari, Oribatida). Soil Organisms 80: 171-195.

Betz, O. (2008), Ökomorphologie: Die Analyse der Form und Funktion von Strukturen in ihrer Beziehung zur Umwelt, Entomologia Generalis 31 (2), 129-139.

Betz, O. & Köhler, H.-R. (Hrsg.) (2008): Die Evolution des Lebendigen. Grundlagen und Aktualität der Evolutionslehre. Attempto, 304 pp.

Betz, O. & Köhler, H.-R. (2008): Die Evolution des Lebendigen: Eine Einführung. In: Betz, O. & Köhler, H.-R. (Hrsg.), Die Evolution des Lebendigen. Grundlagen und Aktualität der Evolutionslehre. Attempto, 9-21.

Betz, O., Koerner, L. & Gorb, S. (2008): Insektenzunge als Vorbild für biphasisch viskose Klebstoffe?, adhäsion 6, 38-41.

Betz, O., Rack, A., Schmitt, C., Ershov, A., Dieterich, A., Körner, L., Haas, D. & Baumbach, T. (2008), High-speed X-ray cineradiography for analyzing complex kinematics in living insects. Synchrotron Radiation News, 21 (5), 34-38.

Heethoff, M. (2008). Molekulare Evolution - Was uns die DNS über unseren Ursprung erzählt. In: Betz, O., Köhler, H. (eds.) Die Evolution des Lebendigen - Grundlagen und Aktualität der Evolutionstheorie. Narr, Franke, Attempto Verlag Tübingen, pp 39-56.

Heethoff, M. (2008) Sex? Nein, danke… labor&more 4/08: 10-12.

Heethoff, M., Cloetens, P. (2008). A comparison of synchrotron X-ray phase contrast tomography and holotomography for non-invasive investigations of the internal anatomy of mites. Soil Organisms 80: 205-215.

Heethoff, M., Helfen, L., Cloetens, P. (2008). Non-invasive 3D-visualization of the internal organization of microarthropods using synchrotron X-ray-tomography with sub-micron resolution. J. Vis. Exp. 15: e737

Laumann, M., Bergmann, P., Heethoff, M. (2008). Some remarks on the cytogenetics of oribatid mites. Soil Organisms 80: 223-232.

Schmelzle, S., Norton, R. A., Helfen, L., Heethoff, M. (2008).The ptychoid defensive mechanism in Euphthiracaroidea (Acari: Oribatida): A comparison of exoskeletal elements. Soil Organisms 80: 233-247.

Schomann, A., Afflerbach, K. & Betz, O. (2008): Predatory behaviour of some Central European pselaphine beetles (Coleoptera: Staphylinidae: Pselaphinae) with remarks on selected morphological features of their heads, European Journal of Entomology, 105, 889-907.

Weide, D. & Betz, O. (2008). Anwendung der Synchrotron Mikro-Röntgentomographie (SR-µCT) in der Insektenmorphologie. Mitteilungen der Deutschen Gesellschaft für allgemeine und angewandte Entomologie 16, 467-472.


Betz, O., Wegst, U., Weide, D., Heethoff, M., Helfen, L., Lee, W.-K., Cloetens, P. (2007), Imaging applications of synchrotron x-ray micro-tomography in biological morphology and biomaterial science. I. General aspects of the technique and its advantages in the analysis of arthropod structure. Journal of Microscopy 227, 51-71.

Heethoff, M., Domes, K., Laumann, M., Maraun, M., Norton, R. A., Scheu, S. (2007). High genetic divergences indicate ancient separation of parthenogenetic lineages of the oribatid mite Platynothrus peltifer (Acari, Oribatida). J. Evol. Biol. 20: 392-402.

Heethoff, M., Koerner, L. (2007). Small but powerful - The oribatid mite Archegozetes longisetosus Aoki (Acari, Oribatida) produces disproportionately high forces. J. Exp. Biol. 210: 3036-3042.

Heethoff, M., Laumann, M., Bergmann, P. (2007) Adding to the reproductive biology of the parthenogenetic oribatid mite Archegozetes longisetosus (Acari, Oribatida, Trhypochthoniidae). Turk. J. Zool. 31: 151-159.

Laumann, M., Norton, R. A.,Weigmann, G., Scheu, S., Maraun, M., Heethoff, M. (2007). Speciation in the parthenogenetic oribatid mite genus Tectocepheus (Acari, Oribatida) as indicated by molecular phylogeny, Pedobiologia 51: 111-122.


Betz, O. (2006): Der Anpassungswert morphologischer Strukturen: Integration von Form, Funktion und Ökologie am Beispiel der Kurzflügelkäfergattung Stenus (Coleoptera, Staphylinidae), Entomologie heute 18, 3-26.

Betz, O. (2006), Ecomorphology: Integration of form, function, and ecology in the analysis of morphological structures, Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie 15, 409-416.

Heethoff, M., Bergmann, P., Norton, R. A. (2006). Karyology and sex determination of oribatid mites. Acarologia 46: 127-131.

Schaefer, I., Domes, K., Heethoff, M., Schneider, K., Schön, I., Norton, R. A., Scheu, S., Maraun, M.. (2006) No evidence for the 'Meselson effect' in parthenogenetic oribatid mites (Oribatida, Acari), J. Evol. Biol. 19: 184-193.


Bauer, T., Talarico, F., Mazzei, A., Giglio, A., Zetto-Brandmayr, T., Brandmayr, P., Betz, O. (2005), Hunting ants in Mediterranean claysoils: life history of Siagona europaea (Coleoptera, Carabidae), Italian Journal of Zoology 72, 33-42.

Lipkow, E. & Betz, O. (2005), Staphylinidae and fungi, Faunistisch-Ökologische Mitteilungen 8, 383-411.


Betz, O. (2004): The eating instruments of spore feeders. German Research 2-3, 27-29.

Betz, O. (2004): Nachgefragt: Wie atmen Maikäfer. natur + kosmos Mai 2004, 72.

Betz, O. & Kölsch, G. (2004), The role of adhesion in prey capture and predator defence in arthropods, Arthropod Structure and Development 33, 3-30.

Lee, W-K., Harrison, J.,  Greenlee, K. et al. (2004), Biomechanics of insect respiration and feeding using synchrotron x-ray imaging, Integrative and Comparative Biology 43, 901-901.


Betz, O. (2003), Structure of the tarsi in some Stenus species (Coleoptera, Staphylinidae): External morphology, ultrastructure, and tarsal secretion, Journal of Morphology 255, 24-43.

Betz, O., Newton, A.F. & Thayer, M.K. (2003), Comparative morphology and evolutionary pathways of the mouthparts of spore-feeding Staphylinoidea (Coleoptera), Acta Zoologica 84 (3), 179-238.

Friis, H., Bauer, T. & Betz, O. (2003), An insect larva with a "pig-snout": structure, and function of the nasale of Hyphydrus ovatus L. (1763) (Coleoptera, Dytiscidae), Journal of Zoology 261, 59-68.

Westneat, M.W., Betz, O., Blob, R.W., Fezzaa, K., Cooper, W.J., Lee, W.-K. (2003), Tracheal respiration in insects visualized with synchrotron X-ray imaging, Science 299, 558-560.


Betz, O. (2002), Performance and adaptive value of tarsal morphology in rove beetles of the genus Stenus (Coleoptera, Staphylinidae), Journal of Experimental Biology 205, 1097-1113.

Lee, W.-K., Fezzaa, K., Westneat, M., Betz, O., Donohue, K. (2002), Imaging applications of synchotron radiation for biomechanics and morphology. Integrative and Comparative Biology 42, 1264-1265.


Betz, O. & Fuhrmann, S. (2001), Life history traits in different life forms of predaceous Stenus beetles (Coleoptera, Staphylinidae), living in waterside environments, Netherlands Journal of Zoology 51 (4), 371-393.

Betz, O. & Mumm, R. (2001), The predatory legs of Philonthus marginatus (Coleoptera, Staphylinidae): Functional morphology and tarsal ultrastructure, Arthropod Structure and Development 30 (2), 77-97.

Betz, O. & Mumm, R. (2001), Die Fangbeine bei Philonthus marginatus (Coleoptera, Staphylinidae): Funktionelle Morphologie und Ultrastruktur der Tarsen, Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie 13, 303-307.

Westneat, M.W., Betz, O., Blob R., Fezzaa, K., Cooper, J., Lee, W.K. (2001), A new view of insect biomechanics: Using synchrotron phase-enhanced x-ray imaging to study insect respiration, locomotion, and feeding, American Zoologist 41 (6), 1623-1624.


Betz, O. (2000), Zum Anpassungswert unterschiedlicher Tarsusformen bei Stenus spp. (Coleoptera, Staphylinidae), Verhandlungen des Westdeutschen Entomologentages 1998, 127-133.

Betz, O. (2000), Beziehungen zwischen Morphologie, Verhalten und Ökologie bei Kurzflügelkäfern der Gattung Stenus(Coleoptera, Staphylinidae), Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie 12, 429-433.

Betz, O., Mumm, R. (2000), The predatory legs of Philonthus marginatus (Coleoptera, Staphylinidae): functional morphology and tarsal ultrastructure, American Zoologist 40 (6): 943-944.


Betz, O. (1999), A behavioural inventory of adult Stenus species (Coleoptera: Staphylinidae), Journal of Natural History 33, 1691-1712.


Betz, O. (1998), Comparative studies on the predatory behaviour of Stenus spp. (Coleoptera: Staphylinidae): the significance of its specialized labial apparatus, Journal of Zoology 244, 527-544.

Betz, O. (1998), Life forms and hunting behaviour of some Central European Stenus species (Coleoptera, Staphylinidae), Applied Soil Ecology 9, 69-74.

Bauer, T., Desender, K., Morwinsky, T. & Betz, O. (1998), Eye morphology reflects habitat demands in three closely related ground beetle species (Coleoptera, Carabidae), Journal of Zoology 245, 467-472.

Kölsch, G. & Betz, O. (1998), Ultrastructure and function of the adhesion-capture apparatus of Stenus species (Coleoptera, Staphylinidae), Zoomorphology 118, 263-272.


Betz, O. (1996), Function and evolution of the adhesion-capture apparatus of Stenus species (Coleoptera, Staphylinidae), Zoomorphology 116, 15-34.


Betz, O. (1995), Vergleichende Untersuchungen zur Bedeutung des Labiums im Beutefangverhalten bei Stenus spp. (Coleoptera, Staphylinidae), Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie 10, 591-594.


Enders, D., Tiebes, J., De Kimpe, N., Keppens, M., Stevens, C., Smagghe, G. & Betz, O. (1993), Enantioselective synthesis and determination of the configuration of stenusine, the spreading agent of the beetle Stenus comma, Journal of Organic Chemistry, 58, 4881-4884.


Betz, O. (1992): Studies on winter-active larvae of the ground beetle Carabus problematicus (Coleoptera, Carabidae). Pedobiologia 36, 159-167.