Publications

Preprints available online

1. Albrecht, J., Bocherens, H., Hobson, K., Drucker, D., Sergiel, A., Swenson, J., Zedrosser, A., Marciszak, A., Iregren, E., Drenzel, L., Kysely, R., Lipecki, G., Makowiecki, D., Wagner, J., Zwijacz-Kozica, T., Fritz, S., Revilla, E., Selva, N., preprint. Trophic adaptation of large terrestrial omnivores to global change. BioRxiv https://doi.org/10.1101/2024.05.07.590891
2. Meleg, I.N., Alberti, F., Drucker, D.G., Naparus-Aljancic, M., Feurdean, A., Robu. M., Vlaicu, M., Naito, Y.I., Boroneant, A., Carciumaru, M., Nitu, E.C., Hofreiter, M., Bocherens, H., Barlow, A., preprint. Ancient biomolecults suggest learned foraging strategy in extinct cave bears. bioRXiv https://doi.org/10.1101/2024.09.23.614392
3. Merkens, L., Anders, T., Traylor, W., Bocherens, H., Hickler, T., preprint. A simple climate-driven semi-mechanistic vegetation model can explain the Pan-Asian extent of the glacial mammoth steppe. Merkens, Lisa and Anders, Tim and Traylor, Wolfgang and Bocherens, Hervé and Hickler, Thomas, A Simple Climate-Driven Semi-Mechanistic Vegetation Model Can Explain the Pan-Asian Extent of the Glacial Mammoth Steppe. Available at SSRN: http://dx.doi.org/10.2139/ssrn.5053507   

Recent peer-reviewed publications (since 2019)

In press

1. Baumann, C., Larbey, C., Ebner, M., Bocherens, H., Hardy, K., in press . The effect of plant food treatment on stable isotopes and their relevance for archaeological studies: A methodological pilot study. Journal of Archaeological Method and Theory https://doi.org/10.1007/s10816-024-09690-5   
2. Ebner, M., Roth-Nebelsick, A., Bocherens, H., Gschwender, F., Baumeister, M., Miranda Ebner, N., Hohberg, K., Gan, H.Y., Schneider, C., Maggs-Kölling, G., Marais, E., Lehmitz, R., Miranda, T., in press. The fog harvesting Namib desert dune grass Stipagrostis sabulicola promotes establishment of oases-ecosystems by modifying substrate and atmosphere conditions.  Journal of Arid Environments
3. Krajcarz, M., Baumann, C., Bocherens, H., Presslee, S., Krajcarz, M.T., in press. Dietary preferences and colla-gen to collagen prey-predator trophic discrimination factors (Δ13C, Δ15N) in Late Pleistocene cave hyena. Quaternary Research https://doi.org/10.1017/qua.2024.43    
4. Suraprasit, K., Yamee, C., Chaimanee, Y., Jaeger, J.-J., Bocherens, H., in press. The Pleistocene grassland-dominated mammal fauna from Tham Kra Duk (Nakhon Si Thammarat, Peninsular Thailand). Historical Biology. https://doi.org/10.1080/08912963.2023.2283936 

2025

1. Bocherens, H., Kirillova, I.V., 2025. Cave lion: Isotopes and dietary paleoecology. Earth History and Biodiversity 3: 100015. https://doi.org/10.1016/j.hisbio.2024.100015 

2024

1. Briatico, G., Bocherens, H., Bonnefille, R., Geraads, D., Melis, R.T., Mussi, M., 2024. A view from the high-lands: the Pleistocene environment between 2 and 0.6 Ma at Melka Kunture (Upper Awasch, Ethiopia), based upon stable isotope analysis. Scientific Reports 14: 6619. https://doi.org/10.1038/s41598-024-56768-x
2. Briatico, G., Mussi, M., Melis, R.T., Geraads, D., Bocherens, H., 2024. Intra-tooth isotopic analysis shows sea-sonal variability in the high-elevation context of Melka Kunture (Upper Awash Valley, Ethiopia). Palaeogeography, Palaeoclimatology, Palaeoecology 655: 112555. https://doi.org/10.1016/j.palaeo.2024.112555   
3. Fordham, D., Brown, S.C., Canteri, E., Austin, J., Lomolino, M., Haythorne, S., Armstrong, E., Bocherens, H., Manica, A., Rey-Iglesia, A., Rahbek, C., Nogues-Bravo, D., Lorenzen, E.D., 2024. 52,000 years of woolly rhinoceros population dynamics reveal extinction mechanism. PNAS 121(24): e2316419121. https://doi.org/10.1073/pnas.2316419121
4. Gan, H.Y., Hohberg, K., Schneider, C., Ebner, M., Marais, E., Miranda, T., Lehmitz, R., Maggs.Kölling, G., Bocherens, H, 2024. The hidden oases: Unveiling trophic dynamics in Namib’s Fog Plant Ecosystems. Scientific Reports 14: 13334. https://doi.org/10.1038/s41598-024-61796-8
5. Hofman-Kaminska, E., Merceron, G., Bocherens, H., Boeskorov, G.G., Egorovich, G.S., Krotova, O.O., Paunovic, M., Protopopov, A.V., Shpansky, A., Kowalczyk, R., 2024. Was the steppe bison a grazing beast in Pleistocene landscapes? Royal Society Open Science 11: 240317 https://doi.org/10.1098/rsos.240317
6. Roditi, E., Bocherens, H., Konidaris, G.E., Athanassiou, A., Tourloukis, V., Karkanas, P., Panagopoulou, E., Harvati, K., 2024. Life history of Palaeoloxodon antiquus reveals Middle Pleistocene glacial refugium in the Mega-lopolis basin, Greece. Scientific Reports  https://doi.org/10.1038/s41598-024-51592-9
7. Scheifler, N., Messineo, P., Bocherens, H., Politis, G., 2024. Isotopic evidence of diet breadth hunter-gatherers changes during the Holocene in the Central Pampean Dunefields (Argentina, South America). American Journal of Biological Anthropology 185: e25039. https://doi.org/10.1002/ajpa.25039  
8. Slimak, L., Vimala, T., Seguin-Orlando, A., Metz, L., Zanolli, C., Joannes-Boyau, R., Frouin, M., Arnold, L.J., Demuro, M., Deviese, T., Comeskey, D., Buckley, M., Camus, H., Muth, X., Lewis, J.E., Bocherens, H., Yvorra, P., Tenailleau, C., Duployer, B., Coqueugniot, H., Dutour, O., Higham, T., Sikora, M., 2024. Long genetic and social isolation in Neanderthals before extinction. Cell Genomics. https://doi.org/10.1016/j.xgen.2024.100593
9. Suraprasit, K., Shoocongdej, R., Wattanapituksakul, A., Chintakanon, K., Bocherens, H., 2024. The late occurrence of specialized hunter-gatherer occupation of tropical rainforests in Pang Mapha, northwestern Thailand. Quaternary Science Reviews 329: 108573. https://doi.org/10.1016/j.quascirev.2024.108573
10. Yravedra, J., Courtenay, L.A., Gutiérrez-Rodriguez, M., Saarinan, J., Égüez, N., Luzón, C., Rodriguez-Alba, J.J., Solano, J.A., Titton, S., Montilla-Jiménez, E., Cámara-Donoso, J., Herranz-Rodriogo, H., Estaca, V., Serrano-Ramos, A., Amorós, G., Azanza, B., Bocherens, H., DeMiguel, D., Fagoaga, M., García-Alix, A., Jiménez-Espejo, F., Kaakinen, A., Munuera, M., Ochando, J., Piñero, P., Sánchez-Bandera, C., Viranta-Kovanen, S., Fortelius, M., Agusti, J., Blain, H.-A., Carrión, J., Barsky, D., Oms, O., Mallol, C., Jiménez-Arenas, J.M., 2024. Not seen before. Unveiling depositional context and Mammuthus meridionalis exploitation at Fuente Nueva 3 (Orce, Southern Iberia) through taxonomy and micromorphology. Quaternary Science Reviews 329: 108561. https://doi.org/10.1016/j.quascirev.2024.108561 

2023

1. Baumann, C., Hussain, S.T., Roblíčková, M., Riede, F., Mannino, M., Bocherens, H., 2023. Earliest evidence for hunter-gatherer impacts on raven diet and ecology in the Early Gravettian of Southern Moravia. Nature Ecology & Evolution 7 : 1302-1314. https://www.nature.com/articles/s41559-023-02107-8
2. Briatico, G., Bocherens, H., Bonnefille, R., Geraads, D., Mussi, M., 2023. Isotopic insights into the the Early Acheulean (1.95 Ma – 1.66 Ma) high elevation paleoenvironments at Melka Kunture (Upper Awash, Ethiopia). Archaeological and Anthropological Sciences 15: 184. https://doi.org/10.1007/s12520-023-01879-1
3. Briatico, G., Bocherens, H., 2023. Middle Pleistocene paleoecology in central Italy. New isotopic insights from fauna tooth enamel of Casal de’ Pazzi (Rome, Italy). Journal of Mediterranean Earth Sciences 15: 141-152 https://doi.org/10.13133/2280-6148/18031
4. Briatico, G., Gioia, P., Bocherens, H., 2023. Diet and habitat of the late Middle Pleistocene mammals from Casal de’ Pazzi site (Rome, Italy) using stable carbon and oxygen isotope ratios. Quaternary International 676: 53-62. https://doi.org/10.1016/j.quaint.2023.11.002
5. Cucchi, T., Harbers, H., Neaux, D., Balasse, M., Garbé, L., Fiorillo, D., Bocherens, H., Drucker, D.G., Za-nolli, C., Cornette, R., Arbogast, R.M., Bréhard, S., Bridault, A., Gourichon, L., Guilaine, J., Manen, C., Perrin, T., Schafberg, R., Tresset, A., Vigne, J.D., Herrel, A., 2023. Phenotypic evolution in wild boars (Sus scrofa) during 4500 years of Neolithic transition in Western Europe. Quaternary Science Reviews 309: 108100 https://doi.org/10.1016/j.quascirev.2023.108100
6. Gimranov, D., Bocherens, H., Kavcik-Graumann, N., Nagel, D., Rabeder, G., 2023. The cave bears from Imanay (Southern Urals, Russia). Historical Biology 35: 580-588. https://doi.org/10.1080/08912963.2022.2056837
7. Gomes, V.S., Lessa, C.M.B., Ribeira de Oliveira, G., Machado Bantim, R.A., Sayão, J., Bocherens, H., Araújo, H.I. Jr, Dantas, M.A.T., 2023. Seasonal variations in diet (δ¹³C) and climate (δ¹⁸O) inferred through toxodonts enamel teeth during the Late Pleistocene in the Brazilian Intertropical Region. Journal of South American Earth Sciences 121: 104148. https://doi.org/10.1016/j.jsames.2022.104148
8. Habinger, S.G., Chavasseau, O., Ducrocq, S., Chaimanee, Y., Jeager, J.J., Sein, C., Aung Naing Soe, Stern, S., Bocherens, H., 2023. Isotopic niche modelling of the mammal fauna from the Pondaung Fm., Myanmar, shows microhabitat differences. Insights into paleoecology and early anthropoid primate habitats. Frontiers in Ecology and Evolution 11:1110331. https://doi.org/103389/fevo.2023.1110331
9. Krajcarz, M.T., Baca, M., Baumann, C., Bocherens, H., Goslar, T., Popović, D., Sudoł-Procyk, M,, Krajcarz, M., 2023. New insights into Late Pleistocene cave hyena chronology and population history – the case of Perspektywiczna Cave, Poland. Radiocarbon 65: 1038-1056. https://doi.org/10.1017/RDC.2023.89
10. Ma, J., Sun, B., Bocherens, H., Deng, T., 2023. New insights on the Plio-Pleistocene Equidae palaeoecology and evolution in East Asia. Palaeogeography, Palaeoclimatology, Palaeoecology 614: 111416. https://doi.org/10.1016/j.palaeo.2023.111416
11. Mussi, M., Méndez-Quintas, E., Barboni, D., Bocherens, H., Bonnefille, R., Briatico, G., Geraads, D., Melis, R.T., Panera, J., Piolo, P., Domínguez, A.S., Rubio Jara, S., 2023. A surge in obsidian exploitation more than 1.2 million years ago at Simbiro III (Melka Kunture, Upper Awash, Ethiopia). Nature Ecology and Evolution  7: 337-346. https://doi.org/10.1038/s41559-022-01970-1    
12. Mussi, M., Skinner, M.M., Melis, R.T., Panera, J., Rubio Jara, S., Davies, T.W., Imbrasas, M., Geraads, D., Bocherens, H., Briatico, G., Le Cabec, A., Hublin, J.-J., Gidna, A., Bonnefille, R., Di Bianco, L., Méndez-Quintas, E., 2023. Early Homo erectus lived at high altitudes and produced both Oldowan and Acheulean tools. Science 382(6671): 713-718. https://www.science.org/doi/10.1126/science.add9115
13. Posth, C., Yu, H., Ghalichi, A., Rougier, H., Crevecoeur, I., Huang, Y., Ringbauer, H., Rohrlach, A.B., Nägele, K., Villalba-Mouco, V., Radzeviciute, R., Ferraz, T., Stoessel, A., Tukhobatova, R., Drucker, D.G., Lari, M., Modi, A., Saupe, T., Scheib, C.L., Catalano, G., Pagani, L., Talamo, S., Fewlass, H., Klaric, L., Morala, A., Rué, M., Madelaine, S., Crépin, L., Caverne, J.B., Bocaege, E., Ricci, S., Boschin, F., Bayle, P., Maureille, B., Le Brun-Ricalens, F., Bordes, J.G., Oxilia, G., Bortolini, E., Bignon-Lau, O., Debout, G., Orliac, M., Zazzo, A., Sparacello, V., Starnini, E., Sineo, L., van der Plicht, H., Pecqueur, L, Merceron, G., Garcia, G., Leuvrey, J.M., Garcia, C.B., Gómez.Olivencia, A., Poltowicz-Bobak, D., Le Luyer, M., Storm, P., Hoffmann, C., Kabacinski, J., Filimonova, T., Shnaider, S., Berezina, N., González-Rabanal, B., González Morales, M.R., Marín-Arroyo, A.B., López, B., Alonso-Llamazares, C., Ronchitelli, A., Polet, C., Jadin, I., Cauwe, N., Soler, J., Coromina, N., Rufi, I., Cottiaux, R., Clark, G., Straus, L.G., Julien, M.A., Renhart, S., Talaa, D., Benazzi, S., Romandini, M., Amkreutz, L, Bocherens, H., Wißing, C., Villotte, S., Fernández-López de Pablo, J., Gómez-Puche, M., Esquembre-Bebia, M.A., Bodu, P., Smits, L., Souffi, B., Jankauskas, R., Kozakaité, J., Cupillard, C., Benthien, H., Wehrberger, K., Schmitz, R.W., Feine, S.C., Schüler, T., Thevenet, C., Grigorescu, D., Lüth, F., Kotula, A., Piezonka, H., Schopper, F., Svoboda, J., Sázelová, S., Chizhevsky, A., Khokhlov, A., Conard, N.J., Valentin, F., Harvati, K., Semal, P., Jungklaus, B., Suvorov, A., Schulting, R., Moiseyev, V., Mannermaa, K., Buzhilova, A., Terberger, T., Caramelli, D., Altena, E., Haak, W., Krause, J., 2023. Paleogenomics of Upper Paleolithic to Neolithic European hunter-gatherers. Nature 615: 117-126.  https://doi.org/10.1038/s41586-023-05726-0
14. Sánchez-Bandera, C., Moreno, A.F., Serrano-Ramos, A., Solano-García, J., Barsky, D., DeMiguel, D., Ochando, J., Saarinen, J., Piñero, P., Lozano-Fernández, I., Courtenay, L.A., Titton, S., Luzón, C., Bocherens, H., Yravedra, J., Fortelius, M., Agustí, J., Carrión, J.S., Oms, O., Blain, H.A., Jiménez-Arenas, J.M., 2023. Glacial/interglacial climate variability in Southern Spain during the late Early Pleistocene and climate backdrop for early Homo in Europe. Palaeogeography, Palaeoclimatology, Palaeoecology 625: 111688. https://doi.org/10.1016/j.palaeo.2023.111688
15. Scheifler, N.A., Merino, M., Vitale, P., Kaufmann, C., Messineo, P., Álvarez, M.C., Bocherens, H., 2023. Isotopic ecology in modern and Holocene populations of Pampas deer (Ozotoceros bezoarticus) from central-eastern Argentina. Implications for conservation biology and ecological models of hunter-gatherer subsistence. Environmental Archaeology: The Journal of Human Palaeoecology 28(2): 45-61.  https://doi.org/10.1080/14614103.2020.1846451  

2022

1. Bergström, A., Stanton, D., Taron, U.H., Frantz, L., Sinding, M.-H.S., Ersmark, E., Pfrengle, S., Cassatt-Johnstone, M., Lebrasseur, O., Girdland-Flink, L., Fernandes, D., Ollivier, Speidel, L., M., Gopalakrishnan, S., Westbury, M., Ramos-Madrigal, J., Feuerborn, T.R., Reiter, E., Gretzinger, J., Münzel, S.C., Swali, P., Conard, N., Carøe, C., Haile, J., Linderholm, A., Androsov, S., Barned, I., Baumann, C., Benecke, N., Bocherens, H., Brace, S., Carden, R.F., Drucker, D.G., Fedorov, S., Gasparik, M., Germonpré, M., Grigoriev, S., Groves, P., Hertwig, S.T., Ivanova, V.V., Janssens, L., Jennings, R.P., Kasparov, A.K., Kirillova, I.V., Kurmaniyazov, I., Kuzmin, Y., Kosintsev, P.A., Lázničková-Galetová, M., Leduc, C., Nikolskiy, P., Nussbaumer, M., O'Drisceoil, C., Orlando, L., Outram, A., Pavlova, E.Y., Perri, A.R., Pilot, M., Pitulko, V.V., Plotnikov, V., Protopopov, A., Rehazek, A., Sablin, M., Seguin-Orlando. A., Storå, J., Verjux, C., Zaibert, V.F., Zazula, G., Crombé, P.M., Hansen, A., Willerslev, E., Leonard, J.A., Götherström, A., Pinhasi, R., Schuenemann, V.J., Hofreiter, M., Gilbert, M.T.P., Shapiro, B., Larson, G., Krause, J., Dalén, L., Skoglund, P., 2022. Gray wolf genomic history reveals a dual ancestry of dogs. Nature 607: 313-320. https://doi.org/10.1038/s41586-022-04824-9
2. Habinger, S., Chavasseau, O., Jaeger, J.J., Chaimanee, Y., Merceron, G., Chit Sein, Aung Naing Soe, Bocherens, H., 2022. Evolutionary ecology of Miocene hominoid primates in Southeast Asia. Scientific Reports 12: 11841. https://doi.org/10.1038/s41598-022-15574-z
3. Hardy, K., Bocherens, H., Miller, J.B., Copeland, L., 2022. Reconstructing Neanderthal diets; the case for plant carbohydrates. Journal of Human Evolution 162: 103105. https://doi.org/10.1016/j.jhevol.2021.103105
4. Kosintsev, P.A., Bocherens, H., Kirillova, I.V., Levchenko, V.A., Zazovskaya, E.P., Shidlovskiy, F.K., Trofimova, S.S., Lindqvist, C., 2022. Paleoecological and genetic analysis of Late Pleistocene bears in Asiatic Russia. Boreas 51(2): 465-480. https://doi.org/10.1111/bor.12570   
5. Krajcarz, M., Van Neer, W., Krajcarz, M.T., Popovic, D., Baca, M., De Cupere, B., Goffette, Q., Küchelmann, H.C., Grezak, A., Iwaszczuk, U., Ottoni, C., Van de Vijver, K., Wilczynski, J., Mulczyk, A., Wiejacki, J., Makowiecki, D., Bocherens, H., 2022. Stable isotopes unveil one millennium of domestic cat paleoecology in Europe. Scientific Reports 12: 12775. https://doi.org/10.1038/s41598-022-16969-8   
6. Milton, E., Stansell, N.D., Bocherens, H., Brownlee, A.H., Chala-Aldana, D., Rademaker, K., 2022. Examining Surface Water δ¹⁸O and δ²H Values in the Western Central Andes: A Watershed Moment for Anthropological Mobility Studies. Journal of Archaeological Sciences 146: 105655. https://doi.org/10.1016/j.jas.2022.105655
7. Muñiz Guinea, F., Cáceres Puro, L.M., Díaz-Zorita Bonilla, M., Vargas Jiménez, J.M., Bocherens, H., Nebelsick, J.H., Donaire Romero, T., Rodriguez-Vidal, J., 2022. Environmental implications and chalcolithic orna-mental use of marine barnacle shells present in the tholos of “La Pastora” (Valencina de la Concepción, Sevilla, Spain). Environmental Archaeology: The Journal of Human Palaeoecology 27(5): 496-507. https://doi.org/10.1080/14614103.2021.1891813
8. Naito, Y.I., Hirose, M., Belmaker, M., Henry, D.O., Osawa, M., Nakazawa, T., Habinger, S., Tung, P., Bocherens, H., Massadeh, S., Kadowaki, S., 2022. Paleoenvironment and human hunting activity during MIS 2 in southern Jordan: Isotope records of prey remains and paleosols. Quaternary Science Reviews 282: 107432. https://doi.org/10.1016/j.quascirev.2022.107432
9. Ochando, J., Carrión, J.S., Altolaguirre, Y., Munuera, M., Amorós, G., Jiménez-Moreno, G., Solano-García, J., Barsky, D., Luzón, C., Sánchez-Bandera, C., Toro-Moyano, I., Saarinen, J., Blain, H.A., Serrano-Ramos, A., Bocherens, H., Oms, O., Agustí, J., Fortelius, M., Jiménez-Arenas, J.M., 2022. Palynological investigations in the Orce archaeological zone, Early Pleistocene of southern Spain. Review of Palaeobotany and Palynology 304: 104725. https://doi.org/10.1016/j.revpalbo.2022.104725
10. Saarinen, J., Fortelius, M., Bocherens, H., Oriol, O., Carrion, J., Agusti, J., Jimenez-Arenas, J.M., 2022. Revisiting the paleoenvironment reconstruction of the Orce sites and the early Homo habitats in western Europe. A response to Palmqvist et al. (2022). Quaternary Science Reviews 297: 107730. https://doi.org/10.1016/j.quascirev.2022.107730   

2021

1. Baumann, C., Pfrengle, S., Münzel, S.C., Thalmann, O., Molak, M., Breidenstein, A., Albrecht, G., Kind, C.-J., Conard, N.J., Drucker, D.G., Verjux, C., Leduc, C., Feuerborn, T., Reiter, E., Giemsch, L., Krause, J., Bocherens, H., Schuenemann, V.J., 2021. A refined proposal for the origin of dogs - The case study of Gnirshöhle, a Magdalenian cave site. Scientific Reports 11: 5137. https://doi.org/10.1038/s41598-021-83719-7
2. Germonpré, M., Lázničková-Galetová, M., Sablin, M., Bocherens, H., Prassack, K., 2021. Some comments on “Friend of Foe? Large canid remains from Pavlovian sites and their archaeozoological context”, a paper by Wilczynski et al. (2020). Journal of Anthropological Archaeology 63: 101329.  https://doi.org/10.1016/j.jaa.2021.101329
3. Germonpré, M., Van den Broeck, M., Lázničková-Galetová, M., Sablin, M., Bocherens, H., 2021. Mothering the orphaned pup, or becoming a dog in the Upper Palaeolithic. Human Ecology: An Interdisciplinary Journal 49(6): 677-689.  https://doi.org/10.1007/s10745-021-00234-z
4. Ma, J., Wang, Y., Baryshnikov, G., Drucker, D.G., Zhang, H., Bocherens, H., Hu, Y., 2021. The Mammuthus-Coelodonta Fauna Complex at its southeastern limit: a biogeochemical paleoecology investigation in Northeast Asia. Quaternary International 591: 93-106. https://doi.org/10.1016/j.quaint.2020.12.024
5. Ramos-Madrigal, J., Sinding, M.H.S., Niemann, J., Samaniego Castruita, J.A., Carøe, C. Mak, S.S.T., Ger-monpré, M., Fedovov, S., Bocherens, H., Pitulko, V.V., Pavlova, E.Y., Nikolskiy, P.A., Kasparov, A.K., Ivanova, V.V., Larson, G., Frantz, L.A.F., Willerslev, E., Meldgaard, M., Feuerborn, T., Petersen, B., Sicheritz-Ponten, T., Bachmann, Wiig, Ø., Hansen, A.J., Gilbert, M.T., Gopalakrishnan, S., 2021. Genomes of Pleistocene Siberian wolves uncover multiple extinct wolf lineages. Current Biology 31: 198-206. https://doi.org/10.1016/j.cub.2020.10.002
6. Rey-Iglesia, A., Lister, A.M., Stuart, A.J., Bocherens, H., Szpak, P., Willerslev, E., Lorenzen, E.D., 2021. Late Pleistocene palaeoecology and phylogeography of woolly rhinoceroses. Quaternary Science Reviews 263: 106993. https://doi.org/10.1016/j.quascirev.2021.106993
7. Saarinen, J., Oksanen, O., Žliobaitė, I., de Miguel, D., Azanza, B., Bocherens, H., Luzón, C., Yravedra, J., Courtenay, L., Blain, H.-A., Sánchez-Bandera, C., Serrano-Ramos, A., Barsky, D., Solano-García, J., Tallavaara, M., Jiménez-Arenas, J.M., Oms, O., Agusti, J., Fortelius, M., 2021. Pliocene to Middle Pleistocene history of climate and environmental changes in the Guadix-Baza Basin and the environmental conditions of early human dispersal into Europe. Quaternary Science Reviews 268: 107132.  https://doi.org/10.1016/j.quascirev.2021.107132
8. Suraprasit, K., Shoocongdej, R., Chintakanon, K., Bocherens, H., 2021. Late Pleistocene human ecology in a savanna ecosystem and its subsequent rainforest occupation in highland northwestern Thailand. Scientific Reports 11:16756 https://doi.org/10.1038/s41598-021-96260-4   

2020

1. Baumann, C., Bocherens, H., Drucker, D.G., Conard, N.J., 2020. Fox dietary ecology as a tracer of human impact on Pleistocene ecosystems. PLoS ONE 15(7): e0235692. https://doi.org/10.1371/journal.pone.0235692    
2. Baumann, C., Starkovich, B.M., Drucker, D.G., Münzel, S.C., Conard, N.C., Bocherens, H. 2020. Dietary niche partitioning among Magdalenian canids in southwestern Germany and Switzerland. Quaternary Science Reviews 227: 106032. https://doi.org/10.1016/j.quascirev.2019.106032
3. Dantas, M.A.T., Cherkinsky, A., Lessa, C.M.F., Santos, L.V., Cozzuol, M.A., Omena, É.C., Lopes da Silva, J.L., Sial, A.N., ∫., 2020. Isotopic paleoecology (δ¹³C, δ¹⁸O) of a Late Pleistocene vertebrate community from Sergipe, NE Brazil. Revista Brasileira de Paleontologia 23(2): 138-152. https://doi.10.4072/rbp.2020.2.05
4. Habinger, S., De Cupere, B., Dövener, F., Pucher, E., Bocherens. H., 2020. Mobility and origin of camels in the Roman Empire through serial stable carbon and oxygen isotopes variations in tooth enamel. Quaternary International 567: 80-91. https://doi.org/10.1016/j.quaint.2020.05.029
5. Krajcarz, M., Krajcarz, M.T., Baca, M., Baumann. C., van Neer, W., Popović, D., Sudoł-Procyk, M., Wach, B., Wilczyński, J., Wojenka, M., Bocherens, H., 2020. Ancestors of domestic cats in Neolithic Central Europe – isotopic evidence of a synanthropic diet. PNAS 117(30): 17710-17719. https://doi.org/10.1073/pnas.1918884117
6. Lipson, M., Ribot, I., Mallick, S., Rohland, N., Olalde, I., Adamski, N., Broomandkhoshbacht, N., Lawson, A.M., Lopez, S., Oppenheimer, J., Stewardson, K., Asombang, R., Bocherens, H., Bradman, N., Cornelissen, E., Crevecoeur, I., Lavachery, P., de Maret, P., Mindzie, C.M., Orban, R., Sawchuk, E., Semal, P., Patterson, N., Hellenthal, G., Lalueza-Fox, C., MacEachern, S., Prendergast, M., Reich, D., 2020. Ancient West African foragers in the context of deep human genetic history. Nature 577: 665-670. https://doi.org/10.1038/s41586-020-1929-1
7. Lister, A., Rivals, F., Barkai, R., Bocherens, H., 2020. Current research on mammoths and their relatives. Quaternary Science Reviews 231: 106174. https://doi.org/10.1016/j.quascirev.2020.106174
8. Loog, L., Thalmann, O., Sinding, M.-H.S., Schuenemann, V., Perri, A., Germonpré, M., Bocherens, H., Witt, K.E., Samaniego Castruita, J.A., Velasco, M.S., Lundstrøm, I.K.C., Wales, N., Sonet, G.E., Frantz, L., Schroeder, H., Budd, J., Jimenez, E.-L.., Fedorov, S., Gasparyan, B., Kandel, A.W., Lázničková-Galetová, M., Napierala, H., Uerpmann, H.-P., Nikolskiy, P.A., Pavlova, E.Y., Pitulko, V.V., Herzog, K.-H., Malhi, R.S., Willer-slev, E., Hansen, A.J., Dobney, K., Gilbert, T.P., Krause, J., Larson G., Eriksson, A., Manica, A., 2020. Modern wolves trace their ancestry to a late Pleistocene expansion from Beringia. Molecular Ecology 162: 2015-2025. https://doi.org/10.1111/mec.15329
9. Lord, E., Dussex, N., Kierczak, M., Díez-del-Molino, D., Ryder, O.A., Stanton, D.W.G., Gilbert, M.T.P., Sánchez-Barreiro F., Zhang G., Sinding, M.H.S., Lorenzen E.D., Fedorov S., Bocherens, H., Nathan, S., Goosens, B., van der Plicht, J., Chan, Y.L., Prost, S., Potapova O., Kirillova, I., Lister, A.M., Heintzman, P.D., Kapp, J., Shapiro, B., Vartanyan, S., Götherström, A., Dalén, L., 2020. Pre-extinction demographic stability and genomic signatures of adaptation in the woolly rhinoceros. Current Biology 30(19): 3871-3879. https://doi.org/10.1016/j.cub.2020.07.046
10. Metz, J., Lampei, C., Bäumler, L., Bocherens, H., Dittberner, H., Henneberg, L., de Meaux, J., Tielbörger, K., 2020. Rapid adaptive evolution to drought in a subset of traits in a multi-site climate change experiment. Ecology Letters 20: 1643-1653. https://doi.org/10.1111/ele.13596
11. Naito, Y., Meleg, I., Robu, M., Drucker, D.G., Wißing, C., Hofreiter, M., Barlow, A., Bocherens, H., 2020. Heavy reliance on plants for Romanian cave bears evidenced by amino acid nitrogen isotope analysis. Scientific Reports 10: 6612. https://doi.org/10.1038/s41598-020-62990-0
12. Oinonen, M., Alenius, T., Arppe, L., Bocherens, H., Etu-Sihvola, H., Helama, S., Huhtamaa, H., Lahtinen, M., Mannermaa, K., Onkamo, P., Vanhanen, S., Wessman, A., 2020. Burdened by nature, buried in water – Resilience carried the Iron Age people through Fimbulvinter. PLoS ONE 15(4): e0231787. https://doi.org/10.1371/journal.pone.0231787    
13. Owocki, K., Kremer, B., Cotte, M., Bocherens, H., 2020. Diet preferences and climate inferred from Oxygen and Carbon isotopes of tooth enamel of Tarbosaurus baatar (Nemegt Formation, Mongolia). Palaeogeography, Palaeoclimatology, Palaeoecology 537: 109190. https://doi.org/10.1016/j.palaeo.2019.05.012
14. Pushkina, D., Saarinen, J., Ziegler, R., Bocherens, H., 2020. Stable isotopic and mesowear reconstructions of paleodiet and habitat of the Middle and Late Pleistocene mammals in South-Western Germany. Quaternary Science Reviews 227: 106026 https://doi.org/10.1016/j.quascirev.2019.106026
15. Ring, S.J., Bocherens, H., Wings, O., Rabi, M., 2020. Divergent mammalian body size in a stable Eocene greenhouse climate. Scientific Reports 10:3987.  https://doi.org/10.1038/s41598-020-60379-7
16. Rufí, I., Drucker, D.G., Bocherens, H., Lloveras, L., Soler, J., Soler, N., 2020. Revision on the occurrence of musk ox (Ovibos moschatus Zimmerman 1780) from the Gravettian of Arbreda Cave (Serinyà, Northeast of Iberian Peninsula): New insights for the study of Iberian cold-adapted faunas. Boreas 49: 858-872. https://doi.org/10.1111/bor.12455
17. Suraprasit, K., Jaeger, J.-J., Shoocongdej, R., Chaimanee, Y., Wattanapituksakul, A., Bocherens, H., 2020. Long-term isotope evidence on the diet and habitat breadth of Pleistocene to Holocene caprines in Thailand: implications for the extirpation and conservation of Himalayan gorals. Frontiers in Ecology and Evolution 8:67. https://doi.org/10.3389/fevo.2020.00067  

2019

1. Arppe, L., Karhu, J.A., Vartanyan, S., Drucker, D.G., Etu-Sihvola, H., Bocherens, H., 2019. Thriving or surviving? The isotopic record of the Wrangel Island woolly mammoth population. Quaternary Science Reviews 222: 105889. https://doi.org/10.1016/j.quascirev.2019.105889
2. Bocherens, H. 2019. Isotopic insights on cave bear palaeodiet. Historical Biology 31: 410-421. https://doi.org/10.1080/08912963.2018.14465419
3. Bons, P.D., Bauer, C.C., Bocherens, H., de Riese, T., Drucker, D.G., Francken, M., Menéndez, L., Uhl, A., van Milligen, B.P., Wißing, C. 2019. Out of Africa by spontaneous migration waves. PLoS ONE 14(4): e0201998. https://doi.org/10.1371/journal.pone.0201998     
4. Díaz-Zorita Bonilla, M., Aranda Jiménez G., Bocherens, H., Escudero Carillo, J., Sánchez Romero, M., Lozano Medina, Á., Alarcón García, E., Milesi García, L., 2019. Multi-isotopic diet determination of Southeastern Iberian Megalithic populations: The cemeteries of El Barranquete and Panoría. Archaeological and Anthropological Science 11(8): 3681-3698. https://doi.org/10.1007/s12520-018-0769-5
5. Etu-Sihvola, H., Bocherens, H., Drucker, D.G., Junno, A.P., Mannermaa, K., Oinonen, M., Uusitaol, J., Arppe, L., 2019. The dIANA database – isotopic baseline for paleodietary research in the Baltic Sea area. Journal of Archaeological Science: Reports 24: 1003-1013. https://doi.org/10.1016/j.jasrep.2019.03.005
6. Gretzinger, J., Molak, M., Reiter, E., Pfrengle, S., Urban, C., Neukamm, J., Blant, M., Conard, N.J., Cupil-lard, C., Dimitrijević, V., Drucker, D.G., Hofman-Kamińska, E., Kowalczyk, R., Paunovic, M., Krajcarz, M.T., Krajcarz, M., Münzel, S.C., Peresani, M, Romandini, M., Rufí, I., Soler, J., Terlato, G., Krause, J., Bocherens, H., Schuenemann, V.J., 2019. Large-scale mitogenomic analysis of the phylogeography of the Late Pleistocene cave bear. Scientific Reports 9: 10700 https://doi.org/10.1038/s41598-019-47073-z
7. Hofman-Kamińska, E., Bocherens, H., Drucker, D.G., Fyfe, R.M., Gumiński, W., Makowiecki, D., Pacher, M., Piličiauskienė, G., Samojlik, T., Woodbridge, J., Kowalczyk, R., 2019.  Adapt or die – response of large herbivores to environmental changes in Europe during the Holocene. Global Change Biology 25(9): 2915-2930. https://doi.org/10.1111/gcb.14733
8. Krajcarz, M.T., Krajcarz, M., Drucker, D.G., Bocherens, H., 2019. Prey-to-fox isotopic enrichment of ³⁴S in bone collagen: Implications for palaeoecological studies. Rapid Communications in Mass Spectrometry 33: 1311-1317. https://doi.org/10.1002/rcm.8471
9. Ma, J., Wang, Y., Jin, C., Hu, Y., Bocherens, H., 2019. Ecological flexibility and differential survival of Pleistocene Stegodon orientalis and Elephas maximus in mainland southeast Asia revealed by stable isotope (C, O) analysis. Quaternary Science Reviews 212: 33-44. https://doi.org/10.1016/j.quascirev.2019.03.021
10. Rey-Iglesia, A., García-Vázquez, A., Treadaway, E.C., van der Plicht, J., Baryshnikov, G.F., Szpak, P., Bocherens, H., Boeskorov, G.G., Lorenzen, E.D., 2019. Evolutionary history and palaeoecology of Brown bear in North-East Siberia re-examined using ancient DNA and stable isotopes from skeletal remains. Scientific Reports  9 : 4462. https://doi.org/10.1038/s41598-019-40168-7
11. Smith, O., Dunshea, G.J., Sinding, M.H.S, Fedorov, S., Germonpre, M., Bocherens, H., Gilbert, M.T.P., 2019. Shotgun sequencing of ancient RNA from late Pleistocene and historical canids shows tissue-specific tran-scriptome survival. PLOS Biology 17(7): e3000166. https://doi.org/10.1371/journal.pbio.3000166    
12. Strani, F., Pushkina, D., Bocherens, H., Bellucci, L., Sardella, R., DeMiguel, D., 2019. Dietary adaptations of Early and Middle Pleistocene fossil equids from the Agnani basin (Frosinone, central Italy). Frontiers in Ecology and Evolution 7: article 176 https://doi:10.3389/fevo.2019.00176      
13. Suraprasit, K., Jongautchariyakul, S., Yamee, C., Pothichaiya, C., Bocherens, H., 2019. New fossil and isotope evidence for the Pleistocene zoogeographic transition and hypothesized savanna corridor in Peninsular Thailand. Quaternary Science Reviews 221: 105861. https://doi.org/10.1016/j.quascirev.2019.105861   
14. Terlato, G., Bocherens, H., Romandini, M., Nannini, N., Hobson, K.A., Peresani, M., 2019. Chronological and isotopic data support a revision for the timing of cave bear extinction in Mediterranean Europe. Historical Biology 31: 474-484. https://doi.org/10.1080/08912963.2018.1448395
15. Wißing, C., Rougier, H., Baumann, C., Comeyne, A., Crevecoeur, I., Drucker, D.G., Gaudzinski-Windheuser, S., Germonpré, M., Gómez-Olivencia, A., Krause, J., Matthies, T., Naito, Y.I., Posth, C., Semal, P., Street, M., Bocherens, H., 2019. Stable isotopes reveal patterns of diet and mobility in last Neandertals and first modern humans in Europe. Scientific Reports 9:4433. https://doi.org/10.1038/s41598-019-41033-3  

Other publications:

1. Bocherens, H., Drucker, D.G., in press. Tomb VII of Qatna: Isotopic investigations on bones and human diet. in Dohmann, H., Degenhardt, S., Lange-Weber, S. und Pfälzner, P. (Eds): Die Gruft VII in Qatna. Befund, Kontex-tualisierung, Interpretation. Qatna Studien 12, Harrassowitz-Verlag, Wiesbaden
2. Starkovich, B.M., Münzel, S.C., Kitagawa, K., Krönneck, P., Riehl, S., Bocherens, H., Drucker, D.G., Conard, N.J., in press. Environment and subsistence during the Aurignacian of the Swabian Jura. Obermaier Conference Heidenheim, Kerns Verlag.
3. Koehler, H., Wegmüller, F., Audiard, B., Auguste, P., Bahain, J.J., Bocherens, H., Diemer, S., Preusser, F., Pümpin, C., Sévêque, N., Stoetzel, E., Tombret, O., Wuscher, P., 2024. The Middle Paleolithic occupations of Mutzig-Rain (Alsace, France). In: (Koehler, H., Conard, N., Floss, H., Lamotte, A, Eds) The Rhine during the Middle Paleolithic: boundary or corridor? Kerns Verlags, Tübingen Publications in Prehistory. Settlement dynamics of the Middle Palaeolithic and Middle Stone Age, volume V. pp. 121-142. https://doi.org/10.51315/9783935751353.006
4. Bocherens, H., Jaouen, K., Rivals, F., Slon, V., Talamo, S., Welker, F., 2022. Chapter 16: Methodological advances in Neanderthal identification, phylogeny, chronology, mobility, climate and diet. In: (F. Rivals, F. Romagnoli, S. Benazzi, Eds) Updating the Neanderthals: understanding behavioural complexity in the Late Middle Palaeolithic,  pp. 303-320.   
5. Rivals, F., Bocherens, H., Camarós, E., Rosell, J., 2022. Diet and ecological interactions in the Middle and Late Pleistocene. In: (F. Rivals, f. Romagnoli, S. Benazzi, Eds) Updating the Neanderthals: understanding behaviour-al complexity in the Late Middle Palaeolithic, pp. 39-54.
6. Bocherens, H., Drucker, D.G., 2021. Isotopic insights on the ecological interactions between humans and woolly mammoths during the Middle and Upper Palaeolithic in Europe. in: Konidaris, G.E., Barkai, R., Tourloukis, V., Harvati, K. (Eds.), Human-elephant interactions: from past to present. Tübingen University Press, Tübingen, pp. 249-262. https://dx.doi.org/10.15496/publikation-55586
7. Germonpré, M., Bocherens, H., Bessudnov, A., Lázničková-Galetová, M., Reynolds, N., Sablin, M., Wißing, C., 2021. Seasonality at Middle and Upper Palaeolithic sites based on the presence and wear of molars from nursing mammoth calves. in: Konidaris, G.E., Barkai, R., Tourloukis, V., Harvati, K. (Eds.), Human-elephant interactions: from past to present. Tübingen University Press, Tübingen, pp. 367-406. https://dx.doi.org/10.15496/publikation-55584   
8. Elalouf, J.-M., Bocherens, H., Bon, C., Fosse, Ph., Palacio, P., Philippe, M., 2020. 6.4. Restes osseux et analyse ADN: apports paléoenvironnementaux. In: Atlas de la grotte Chauvet-Pont d’Arc, Delannoy J.J. & Geneste J.M. (Eds), Collection “Documents d’archéologie française”, Editions Fondation Maison des Sciences de l’Homme, pp. 150-152.
9. Germonpré, M., Lázničková-Galetová, M., Sablin, M., Bocherens, H., 2020. 11. Could incipient dogs have en-hanced differential access to resources among Upper Palaeolithic hunter-gatherers in Europe? In: (L. Moreau, ed.), Social Inequality before farming? Multidisciplinary approaches to the investigation of egalitarian and non-egalitarian social relationships in prehistoric and extant hunter-gatherer societies McDonald institute of Archaeology, Cambrigde, pp. 179-200. e-book with open access. https://www.repository.cam.ac.uk/handle/1810/313518
10. Valladas, H., Bocherens, H., Bon, C., Fosse, Ph., Philippe, M., Quiles, A., Geneste, J.-M., Monney, J., 2020. 5.4. Datation 14C d’ossements. In: Atlas de la grotte Chauvet-Pont d’Arc, Delannoy J.J. & Geneste J.M. (Eds), Collection “Documents d’archéologie française”, Editions Fondation Maison des Sciences de l’Homme, p. 125.

Complete List, from March 2024.

To download publications as pdf documents, please click here. (Academia) or here (Researchgate)