Paläoanthropologie

Publications

FIRSTSTEPS team members and collaborators are highlighted in bold.

2026

Harvati K. 2026. The role of the Eastern Mediterranean in human evolution: Recent results from Greece. Journal of the Royal Anthropological Institute. DOI:10.1111/1467-9655.70138  

Naumann D., Harvati K. 2026. A reinvestigation of the Upper Paleolithic postcranial human remains from the La Rochette rock shelter (Saint-Léon-sur-Vézère, France). PaleoAnthropology 2026:1, 1-27. DOI: 10.48738/2026.iss1.3869

Bosman A., Buck L.T., Reyes-Centeno H., Mirazón Lahr M., Stringer C., Harvati K. 2026. The morphological affinities of the fossil cranium from Kabua, Kenya. Journal of the Royal Anthropological Institute. DOI: 10.1111/1467-9655.70137 

Röding C., Profico A., Merkel M., Harvati K. 2026. A morphological analysis of the modern human frontal bone from Hahnöfersand, Germany. Scientific Reports 16, 12696. DOI: 10.1038/s41598-026-48468-5 

Bludau I.J.E., Wagner B., Kirscher U., Schindler M., Zachow C., Schmidt J., Leicher N., Muttoni G., Perini S., Tourloukis V., Panagopoulou E., Karkanas P., Harvati K., Junginger A. 2026. Drilling the Marathousa paleo-lake in Greece (Peloponnese): Inferring the environmental context of a Middle-Pleistocene archaeological site. Boreas. DOI: 10.1111/bor.70061 

Milks A., Ntinou M., Koutalis S., Michailidis D., Konidaris G., Giusti D., Thompson N., Tsartsidou G., Panagopoulou E., Tourloukis V., Karkanas P., Harvati K. 2026. Evidence for the earliest hominin use of wooden hand-held tools found at Marathousa 1 (Greece). Proceedings of the National Academy of Sciences USA 123 (6), e2515479123. DOI: 10.1073/pnas.2515479123

2025

Kyrikou S., Marinova E., Bludau I.J.E., Karkanas P., Panagopoulou E., Tourloukis V., Junginger A., Harvati K. 2025. The palynological record from Marathousa 1, South Greece, during Marine Isotope Stage (MIS) 12: Local environment and fire history. Quaternary Research 128, 71-90. DOI:  10.1017/qua.2024.59

Konidaris G.E., Aytek A., Kostopoulos D., Yavuz A., Tarhan E., Alçiçek M., Uyar N., Harvati K. 2025. Kayaca, a new vertebrate locality from the Upper Miocene of Türkiye and its importance for the Turolian biogeography of the eastern peri-Mediterranean Region. Palaeobiodiversity and Palaeoenvironment 106, 211-227. DOI:  10.1007/s12549-025-00666-1

De Caro D., Kuhn M., Thompson N., Panagopoulou E., Harvati K., Tourloukis V. 2025. Small flakes for sharp needs: technological behaviour in the Lower Palaeolithic site of Marathousa 1, Greece. PLoS ONE 20(6): e0324958. DOI: 10.1371/journal.pone.0324958

Röding C., El Zaatari S., Ramirez-Rozzi F.V., Stringer C., Burgess M.L., Lacruz R.S., Harvati K. 2025. Dentition of the Mugharet el’Aliya fossil human maxilla, Morocco. American Journal of Biological Anthropology  186, e70015. DOI:  10.1002/ajpa.70015

 Piccirilli E., Sorrentino R., Seghi F., Vazzana A., Belcastro M.G., Talamo S., Harvati K., Bettuzzi M., Morigi M.P., Spagnolo V., Martini I., Moroni A., Boschin F., Ricci S., Benazzi S. 2025. The human remains from the MIS 6 site of Grotta del Poggio (Cilento, southern Italy). A taxonomic and chronological reassessment. American Journal of Biological Anthropology 188, e70188. DOI: 10.1002/ajpa.70188

 Salazar-Fernández A., Carretero J.M., Quintino Y., Harvati K., Rodríguez L., García-González R. 2025. Pre-fusion shape changes of humeral metaphyseal surfaces: a new method for assessing maturity and age in non-adult skeletal individuals. American Journal of Biological Anthropology 186, e25063. DOI: 10.1002/ajpa.25063

2024

Harvati K., Stringer C., Folorunso A. 2024. Comparative 3D shape analysis of the Iwo Eleru mandible, Nigeria. PaleoAnthropology 2024:1, 99-111. DOI: 10.48738/2024.iss1.1084

Roditi E., Bocherens H., Konidaris G.E., Athanassiou A., Tourloukis V., Karkanas P., Panagopoulou E., Harvati K.2024. Life-history of Palaeoloxodon antiquus reveals Middle Pleistocene glacial refugium in the Megalopolis basin, Greece. Scientific Reports 14, 1390. DOI: 10.1038/s41598-024-51592-9

Konidaris G.E., Athanassiou A., Tourloukis V., Chitoglou K., van Kolfschoten T., Giusti D., Thompson N., Tsartsidou G., Roditi E., Panagopoulou E., Karkanas P., Harvati K. 2024. The late Early–Middle Pleistocene mammal fauna from the Megalopolis Basin (Peloponnese, Greece) and its importance for biostratigraphy and palaeoenvironment. Quaternary 7, 41. DOI: 10.3390/quat7040041

Butiseaca G.A., van der Meer M.T.J., Bludau I.J.E., Karkanas P., Vasiliev I., Tourloukis V., Junginger A., Mulch A., Panagopoulou E., Harvati K. 2024. Impact of the MIS 12 glacial stage on the middle Pleistocene hominins in the eastern Mediterranean (Megalopolis basin - Greece). Global and Planetary Change 242, 104585. DOI: 10.1016/j.gloplacha.2024.104585  

Kunze J., Harvati K., Hotz G., Karakostis F.A. 2024. Human-like manual activities in Australopithecus. Journal of Human Evolution 196, 103591. DOI: 10.1016/j.jhevol.2024.103591

Rathmann H., Vizzari M.T., Beier J., Bailey S., Ghirotto S., Harvati K. 2024. Human population dynamics in Upper Paleolithic Europe inferred from fossil dental phenotypes. Science Advances 10, eadn8129. DOI: 10.1126/sciadv.adn8129

Silvestrini S., Real C., Federico Lugli F., Delpiano D., Harvati K., Collina C., Peresani M., Spinapolice E.E., Romandini M., Benazzi S. 2024. Hunting game: new data on the subsistence strategies during the Uluzzian in Italy. Journal of Archaeological Science: Reports 57, 104575. DOI: 10.1016/j.jasrep.2024.104575

Limmer L.S., Santon M., McGrath K., Harvati K., ElZaatari S. 2024. Differences in childhood stress between Neanderthals and early modern humans as reflected by dental enamel growth disruptions. Scientific Reports 14, 11293. DOI: 10.1038/s41598-024-61321-x  

Mori T., Riga A., Aytek A. I., Harvati K. 2024. A new virtual reconstruction and geometric morphometric analysis of the Kocabaş hominin fossil from Turkey. Journal of Human Evolution 191, 103517. DOI: 10.1016/j.jhevol.2024.103517 

Gretzinger J., Schmitt F., Mötsch A., Carlhoff S., Lamnidis T.C., Huang Y., Ringbauer H., Knipper C., Francken M., Mandt F., Hansen L., Freund C., Huang Y., Ringbauer H., Posth C., Rathmann H., Harvati K., Wieland G., Granehaell L., Maixner F., Zink A., Schier W., Krausse D., Krause J., Schiffels S. 2024. Evidence for dynastic succession among early Celtic elites. Nature Human Behaviour 8, 1467-1480. DOI: 10.1038/s41562-024-01888-7

2023

Schmidt, P., Koch T.J., Charrié-Duhaut A., Karakostis, F.A., Harvati, K., Dresely, V., Blessing, M.A. 2023. Production method of the Königsaue birch tar documents cumulative culture in Neanderthals. Journal of Archaeological and Anthropological Science 15, 84. DOI: 10.1007/s12520-023-01789-2

Konidaris G.E., Tourloukis V., Boni G., Athanassiou A., Giusti D., Thompson N., Syrides G., Panagopoulou E., Karkanas P., Harvati K. 2023. Marathousa 2: A New Middle Pleistocene Locality in the Megalopolis Basin (Greece) With Evidence of Hominin Exploitation of Megafauna (Hippopotamus). PaleoAnthropology 2023:1, 34-55 . DOI: 10.48738/2023.iss1.810

Röding C., Stringer C., Lacruz R.S., Harvati K. 2023. Mugharet el’Aliya: affinities of an enigmatic North African Aterian maxillary fragment. American Journal of Biological Anthropology 180(2), 352-369. DOI:  10.1002/ajpa.24642

Rathmann H., Perretti S., Porcu V., Hanihara T., Scott G.R., Irish J.D., Reyes-Centeno H., Ghirotto S., Harvati K. 2023. Inferring human neutral genetic variation from craniodental phenotypes. PNAS Nexus 2, 1-11. DOI: 
10.1093/pnasnexus/pgad217

Posth C., … Harvati K., Semal P., Jungklaus B., Suvorov A., Schulting R., Moiseyev V., Mannermaa K., Buzhilova A., Terberger T., Caramelli D., Altena E., Haak W., Krause J. 2023. Paleogenomics of Upper Paleolithic to Neolithic European hunter-gatherers. Nature 615, 117–126. DOI:  10.1038/s41586-023-05726-0

Buzi C., Profico A., Liang C., Khonsari R.H., O’Higgins P., Moazen M., Harvati K. 2023. Icex: advances in the automatic extraction and volume calculation of cranial cavities. Journal of Anatomy 242(6), 1172-1183. DOI: 10.1111/joa.13843

2022

Harvati K., Reyes Centeno H. 2022. Evolution of Homo in the Middle and Late Pleistocene. Journal of Human Evolution 173, 103279. DOI: 10.1016/j.jhevol.2022.103279

Harvati K., Ackermann R.R. 2022. Merging morphological and genetic evidence to assess hybridization in Western Eurasian Late Pleistocene hominins. Nature Ecology & Evolution 6, 1573-1585. DOI: 10.1038/s41559-022-01875-z

Balzeau A., Albessard-Ball L., Kubicka A.M., Filippo A., Beaudet A., Santos E., Bienvenu T., Arsuaga J.-L., Bartsiokas A., Berger L., Bermúdez de Castro J.M., Brunet M., Carlson K. J., Daura J., Gorgoulis V.G., Grine F.E., Harvati K., Hawks J., Herries A., Hublin J.-J., Hui J., Ives R., Joordens J.A., Kaifu Y., Kouloukoussa M., Léger B., Lordkipanidze D., Margvelashvili A., Martin J., Martinón-Torres M., May H., Mounier A., du Plessis A., Rae T., Röding C., Sanz M., Semal P., Stratford D., Stringer C., Tawane M., Temming H., Tsoukala E., Zilhão J., Zipfel B., Buck L.T. 2022. Frontal sinuses and human evolution. Science Advances 8, eabp9767. DOI: 10.1126/sciadv.abp9767