Paläoanthropologie

Selected Publications

Edited Volumes

Harvati K., Ioannidou M., (Eds.) 2025. Human Evolution at the CROSSROADS, Proceedings of the Closing Symposium, February 2022. Tübingen Paleoanthropology Book Series – Contributions in Paleoanthropology III. Tübingen University Press: Tübingen.

Francken M.Harvati K. (Eds.) 2023. Facets of Osteology: An honorary volume for Prof. Dr. Joachim Wahl, pp. 1-245. In: K. Harvati (Series Editor) Tübingen Paleoanthropology Book Series: Contributions in Paleoanthropology II. Tübingen University Press: Tübingen.

Reyes Centeno H., Harvati K. (Eds.) 2021. Ancient Connections in Eurasia. Words, Bones, Genes, Tools: DFG Centre for Advanced Studies Series. Kerns Verlag, Tübingen.

Konidaris G., Barkai R., Tourloukis V., Harvati K. 2021. Human elephant interactions: from past to present, pp. 1-398. Tübingen Paleoanthropology Book Series: Contributions in Paleoanthropology 1. Tübingen University Press: Tübingen.

Harvati K., Konidaris G., Tourloukis V. (Eds.) 2018. Special Issue Human Evolution at the Gates of Europe. Quaternary International Special Issue Volume 497 Part A, pp. 1-240.

Harvati K., Jäger G., Reyes-Centeno H. (Eds.) 2018. New Perspectives on the Peopling of the Americas. Words, Bones, Genes, Tools: DFG Centre for Advanced Studies Series. Kerns Verlag, Tübingen.

El Zaatari S., Harvati K., Bae C. (Eds.) 2018. Special Issue Filling the Gaps in the Human Evolutionary Story. Quaternary International, Volume 466 Part A, pp. 1-106.

Harvati K., Roksandic, M. (Eds.) 2016. Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Vertebrate Paleobiology and Paleoanthropology Series, Springer: Dordrecht.

Harvati K., Harrison, T. (Eds.) 2006. Neanderthals Revisited: New Approaches and Perspectives. Vertebrate Paleobiology and Paleoanthropology Series, Springer: Dordrecht.

Selected recent journal articles

Harvati K. 2026. The role of the Eastern Mediterranean in human evolution: Recent results from Greece. Journal of the Royal Anthropological Institute https://doi.org/10.1111/1467-9655.70138 

Naumann D., Harvati K. 2026. A reinvestigation of the Upper Paleolithic postcranial human remains from the La Rochette rock shelter (Saint-Léon-sur-Vézère, France). PaleoAnthropology 2026:1, 1-27  https://paleoanthropology.org/ojs/index.php/paleo/article/view/3869/3997

Bosman A., Buck L.T., Reyes-Centeno H., Mirazón Lahr M., Stringer C., Harvati K. 2026. The morphological affinities of the fossil cranium from Kabua, Kenya. Journal of the Royal Anthropological Institute https://doi.org/10.1111/1467-9655.70137 (invited contribution)

Röding C., Profico A., Merkel M., Harvati K. 2026. A morphological analysis of the modern human frontal bone from Hahnöfersand, Germany. Scientific Reports 16, 12696. https://doi.org/10.1038/s41598-026-48468-5

Milks A., Ntinou M., Koutalis S., Michailidis D., Konidaris G., Giusti D., Thompson N., Tsartsidou G., Panagopoulou E., Tourloukis V., Karkanas P., Harvati K. 2026. Evidence for the earliest hominin use of wooden hand-held tools found at Marathousa 1 (Greece). Proceedings of the National Academy of Sciences USA 123 (6), e2515479123b https://doi.org/10.1073/pnas.2515479123

Kyrikou S., Marinova E., Bludau I.J.E., Karkanas P., Panagopoulou E., Tourloukis V., Junginger A., Harvati K. 2025. The palynological record from Marathousa 1, South Greece, during Marine Isotope Stage (MIS) 12: Local environment and fire history. Quaternary Research 128, 71-90  https://doi.org/10.1017/qua.2024.59

De Caro D., Kuhn M., Thompson N., Panagopoulou E., Harvati K., Tourloukis V. 2025. Small flakes for sharp needs: technological behaviour in the Lower Palaeolithic site of Marathousa 1, Greece. PLoS ONE 20(6): e0324958. https://doi.org/10.1371/journal.pone.0324958   OPEN ACCESS

Röding C., El Zaatari S., Ramirez-Rozzi F.V., Stringer C., Burgess M.L., Lacruz R.S., Harvati K. 2025. Dentition of the Mugharet el’Aliya fossil human maxilla, Morocco. American Journal of Biological Anthropology  186, e70015 https://doi.org/10.1002/ajpa.70015

Konidaris G.E., Athanassiou A., Tourloukis V., Chitoglou K., van Kolfschoten T., Giusti D., Thompson N., Tsartsidou G., Roditi E., Panagopoulou E., Karkanas P., Harvati K. 2024. The late Early–Middle Pleistocene mammal fauna from the Megalopolis Basin (Peloponnese, Greece) and its importance for biostratigraphy and palaeoenvironment. Quaternary 7, 41 https://doi.org/10.3390/quat7040041

Butiseaca G.A., van der Meer M.T.J., Bludau I.J.E., Karkanas P., Vasiliev I., Tourloukis V., Junginger A., Mulch A., Panagopoulou E., Harvati K. 2024. Impact of the MIS 12 glacial stage on the middle Pleistocene hominins in the eastern Mediterranean (Megalopolis basin - Greece). Global and Planetary Change 242, 104585 https://doi.org/10.1016/j.gloplacha.2024.104585 

Kunze J.Harvati K., Hotz G., Karakostis F.A. 2024Human-like manual activities in Australopithecus. Journal of Human Evolution 196103591 10.1016/j.jhevol.2024.103591

Rathmann H., Vizzari M.T., Beier J., Bailey S., Ghirotto S., Harvati K. 2024. Human population dynamics in Upper Paleolithic Europe inferred from fossil dental phenotypes. Science Advances 10, eadn8129   DOI: 10.1126/sciadv.adn8129

Mori T., Riga A., Aytek A. I., Harvati K. 2024Virtual reconstruction and geometric morphometric analysis of the Kocabaş hominin fossil from Turkey: Implications for taxonomy and evolutionary significance. Journal of Human Evolution 191, 103517 https://doi.org/10.1016/j.jhevol.2024.103517 

Harvati K., Stringer C., Folorunso A. 2024. Comparative 3D shape analysis of the Iwo Eleru mandible, Nigeria. PaleoAnthropology 2024:1, 99-111 https://doi.org/10.48738/2024.iss1.1084

Roditi E., Bocherens H., Konidaris G.E., Athanassiou A., Tourloukis V., Karkanas P., Panagopoulou E., Harvati K. 2024. Life-history of Palaeoloxodon antiquus reveals Middle Pleistocene glacial refugium in the Megalopolis basin, Greece. Scientific Reports 14, 1390 https://www.nature.com/articles/s41598-024-51592-9

Siali C., Niinimäki S., Harvati K., Karakostis F.A. 2024. Reconstructing patterns of domestication in reindeer using 3D muscle attachment areas. Archaeological and Anthropological Sciences 16, 19. https://doi.org/10.1007/s12520-023-01910-5

Konidaris G.E., Tourloukis V., Boni G., Athanassiou A., Giusti D., Thompson N., Syrides G., Panagopoulou E., Karkanas P., Harvati K. 2023. Marathousa 2: A New Middle Pleistocene Locality in the Megalopolis Basin (Greece) With Evidence of Hominin Exploitation of Megafauna (Hippopotamus). PaleoAnthropology 2023:1,34-55 https://doi.org/10.48738/2023.iss1.810

Röding C., Stringer C., Lacruz R.S., Harvati K. 2023. Mugharet el’Aliya: affinities of an enigmatic North African Aterian maxillary fragment. American Journal of Biological Anthropology 180, 352-369  https://doi.org/10.1002/ajpa.24642

Harvati K., Ackerman R.R. 2022. Merging Morphological and Genetic Evidence to assess hybridization in Western Eurasian Late Pleistocene hominins. Nature Ecology & Evolution 6, 1573-1585 

Harvati K., Reyes Centeno H. 2022. Evolution of Homo in the Middle and Late Pleistocene. Journal of Human Evolution 173, 103279

Guibert-Cardin J., Tourloukis V., Thompson N., Panagopoulou E., Harvati K., Nicoud E., Beyries S. 2022. The function of small tools in Europe during the Middle Pleistocene: The case of Marathousa 1 (Megalopolis, Greece). Journal of Lithic Studies 9(1), 1-29.

Röding C., Stringer C., Lacruz R.S., Harvati K. 2022. Mugharet el’Aliya: affinities of an enigmatic North African Aterian maxillary fragment. American Journal of Biological Anthropology 

Kunze J., Karakostis F.A., Merker S., Peresani M., Hotz G., Tourloukis V., Harvati K. 2022. Entheseal patterns suggest habitual tool use in early hominins. PaleoAnthropology 2022:2, 195-210 

Konidaris G., Athanassiou A., Panagopoulou E., Harvati K. 2022. First record of Macaca (Cercopithecidae, Primates) in the Middle Pleistocene of Greece. Journal of Human Evolution 162, 103104

Karakostis F.A., Reyes-Centeno H., Franken M., Hotz G., Rademaker K., Harvati K. 2021. Bio-cultural evidence of precise manual activities in an Early Holocene individual of the high-altitude Peruvian Andes. American Journal of Physical Anthropology 174, 35-48

Karakostis F.A., Harvati K. 2021. New horizons in reconstructing past human behavior: Introducing the “Tübingen University Validated Entheses-based Reconstruction of Activity” method. Evolutionary Anthropology 30, 185-198

Karakostis F.A., Häufle D., Anastopoulou I., Moraitis K., Hotz, G., Tourloukis V., Harvati K. 2021. Biomechanics of the human thumb and the evolution of dexterity. Current Biology 31, 1-9

McGrath, K, Limmer, LS, Lockey, AL, Guatelli-Steinberg, D, Reid, DJ, Wityel, C, Bocaege, E, McFarlin, SC, El Zaatari, S. 2021. 3D enamel profilometry reveals faster growth but similar stress severity in Neanderthal versus Homo sapiens teeth. Scientific Reports 11:522.

Bosman A., Reyes-Centeno H., Harvati K. 2020. A virtual assessment of the suprainiac depressions on the Eyasi I (Tanzania) and Aduma ADU-VP-1/3 (Ethiopia) Pleistocene hominin crania. Journal of Human Evolution 145, 102815

Rathmann, H., Reyes-Centeno H. 2020. Testing the utility of dental morphological trait combinations for inferring human neutral genetic variation. Proceedings of the National Academy of Sciences (USA) 117(20), 10769-10777

Harvati K., Röding C., Bosman A., Karakostis F.A., Grün R., Stringer C., Karkanas P., Thompson N.C., Koutoulidis V., Moulopoulos L.A., Gorgoulis V.G., Kouloukoussa M. 2019. Apidima Cave fossils provide the earliest evidence of Homo sapiens in Eurasia. Nature 571, 500-504

Mori T., Harvati K. 2019. Basicranial ontogeny comparison in Pan and Homo and its use for developmental stage definition of KNM-ER42700. American Journal of Physical Anthropology 170, 579-594

Lacruz R.S., Stringer C.B., Kimbel W.H., Wood B., Harvati K., O’Higgins P., Bromage T.G., Arsuaga J.-L. 2019. The evolutionary history of the human face. Nature Ecology & Evolution 3, 726-736

Kranioti E., Grigorescu D., Harvati K. 2019. State of the art forensic techniques reveal evidence for interpersonal violence ca. 30.000 years ago. PLoSONE 14(7): e0216718

Francken M., Beier J., Reyes-Centeno H., Harvati K., Rademaker K. 2018. The human skeletal remains from the Cuncaicha rockshelter, Peru. In (K. Harvati, G. Jäger, H. Reyes-Centeno, eds.): New Perspectives on the Peopling of the Americas, pp. 125-152.

Beier J., Anthes N., Wahl J., Harvati K. 2018. Similar cranial trauma prevalence among Neanderthals and Upper Paleolithic humans. Nature 563, 686-690

Karakostis F.A., Hotz G., Tourloukis V., Harvati K. 2018. Evidence for precison grasping in Neandertal daily activities. Science Advances 4, eaat2369

El Zaatari, S. 2018. The central Levantine corridor: the Paleolithic of Lebanon. Quaternary International 466:33-47.

Tourloukis V., Harvati K. 2018. The Palaeolithic record of Greece: a synthesis of the evidence and a research agenda for the future. Quaternary International, 466, 48-65

Posth C., Nakatsuka N., Lazaridis I., Skoglund P., Mallick S., Lamnidis T.C., Rohland N., Nägele K., Adamski N., Bertolini E., Broomandkhoshbacht N., Cooper A., Culleton B.J., Ferraz da Silva T., Ferry M., Furtwängler A., Haak W., Harkins K., Harper T.K., Hünemeier T., Lawson A.M., Llamas B., Michel M., Nelson E., Oppenheimer J., Patterson N., Schiffels S., Sedig J., Stewardson K., Talamo S., Wang C.-C., Hublin J.-J., Hubbe M., Harvati K., Beier J., Francken M., Kaulicke P., Reyes-Centeno H., Rademaker K., Trask W.R., Robinson M., Gutierrez S.M., Prufer K.M., Salazar-Garcia D.C., Chim E.N., Müller Plumm Gomes L., Lopes Alves M., Liryo A., Inglez M., Oliveira R.E., Bernardo D., Barioni A., Wesolowski V., Scheifler N.A., Rivera M.A., Plens C.R., Messineo P., Figuti L., Corach D., Scabuzzo C., Eggers S., Dantas de Blasis P.R., Reindel M., Méndez C., Politis G., Tomasto-Cagigao E., Kennett D.J., Strauss A., Fehren-Schmitz L., Krause J., Reich D. 2018. Reconstructing the deep population history of Central and South America. Cell 175, 1185-1197

Hublin J.J., Ben-Ncer A., Bailey S., Freideline S., Neubauer S., Skinner M.M., Bergmann I., Le Cabec A., Benazzi S., Harvati K., Gunz P. 2017. New fossils from Jebel Irhoud (Morocco) and the Pan-African origin of Homo sapiens. Nature 546, 289-292.

Rathmann H., Reyes-Centeno H., Ghirotto S., Creanza N., Hanihara T., Harvati K. 2017.  Reconstructing human population history from dental phenotypes. Scientific Reports 7, 12495.

Sahle, Y, El Zaatari, S, White, TD. 2017. Hominid butchers and biting crocodiles in the African Plio-Pleistocene. Proceedings of the National Academy of Science, USA 114:13164-13169.

Singh N., Albert F.W., Plyusnina I., Trut L., Pӓӓbo S., Harvati K. 2017. Facial shape differences between rats selected for tame and aggressive behaviors. PLOS ONE 12(4): e0175043

Fu Q., Posth C., Hajdinjak M., Petr M., Mallick S., Fernandes D., Furtwängler A., Haak W., Meyer M., Mittnik A., Nickel B., Peltzer A., Rohland N., Slon V., Lazaridis I., Lipson M., Schiffels S., Skoglund P., Gély B., Benazzi S., Gonzales Morales M.R., Peresani M., Straus L.G., Caramelli D., Lari M., Ronchitelli A., Valentin F., Constantin S., Coppola D., Grigorescu D., Neugebauer-Maresch C., Thevenet C., Wehrberger K., Crevecoeur I., Rougier H., Semal P., Cupillard C., Bocherens H., Mannino M., Conard N.J., Harvati K., Moiseyev V., Drucker D.G., Svoboda J., Pinhasi R., Kelso J., Patterson N., Krause J., Pääbo S., Reich D. 2016. The genetic history of Ice Age Europe. Nature 524, 200-2005

Reyes-Centeno H., Ghirotto S., Détroit F., Grimaud-Hervé D., Barbujani G., Harvati K. 2014. Genomic and Cranial Phenotype Data Support Multiple Modern Human Dispersals from Africa and a Southern Route into Asia. Proceedings of the National Academy of Sciences (USA) 111, 7248-7253